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glabrous and dark shiny green, below finely 
appressed brown pilosulous on the midrib and 
slightly so on the lateral veins, the outer three- 
quarters coarsely remotely crenate serrate, the 
lateral veins 8-17 on a side, straight, ascending, 
then near the margin the tips inarching and 
branches salient in the serrae; cymes 1 -flowered, 
from the leafy axils, sparsely brown pilose; 
peduncle 2-3 mm long, diverging in flower; 
pedicels 2-6 mm long in flower, 7-14 mm long 
in fruit, sparsely brown pilose; the paired bracts 
of the pedicel 2.5-3 mm long, caducous, lan- 
ceolate, brown subappressed pilose, sparsely so 
on the back, densely so near the margins; buds 
26-27 mm long when fresh, without very 
remotely brown pilose, within closely appressed 
brown pilose, fusiform, but the body oblanceo- 
loid, the beak 8-10 mm long, ascending and 
arcuate or subsigmoid, the 5 lobes almost com- 
pletely fused, but at apex of the upper lip there 
are 3 minute tips like rounded nipples, and 2 
shorter ones of the lower lip, the body and 
lower part of the beak at anthesis opening by a 
distal slit 10 mm long, through which the 
corolla protrudes; calyx 25-27 mm long, cadu- 
cous from the young developing fruit; corolla 
28 mm long when fresh, white, glabrous with- 
out, the tube 20 mm long, glabrous except for the 
throat which is capitate glandular puberulent, 
cuneate-cylindric, but at mid-point deflected at 
40° from the axis of the lower tube, at base 
3 mm in diameter, at the middle 5 mm, at the 
throat 8 mm high and 5 mm wide, limb 2- 
lipped, 5-lobed, upper lip spreading at 80° to 
the axis of the throat, the two lobes 6 mm long, 
7 mm wide, suborbicular, within capitate glan- 
dular puberulent on lower half; lower lip 3- 
lobed; the two lateral lobes 8 mm long, 8.5 mm 
wide, oblong suborbicular, with similar glan- 
dular puberulence; lower lobe 9 mm long, 8.5 
mm wide, broadly elliptic, the upper surface 
capitate glandular except near the tip and below; 
the two lower stamens perfect, their filaments 
adnate to the corolla tube to within 4 mm of 
the throat, the free portion 2.5 mm long, 
stout, spirally upcurved; the two perfect anthers 
3.5 mm long, connivent, obliquely and asym- 
metrically broad ovate, the connective similar 
in shape; the two lateral staminodia adnate to 
the corolla to within 6 mm of the throat, the 
free filament tips 0.3 mm long, bearing an 
antheroid tip 0.7 mm long, translucent; the 
upper staminodium similar but attached 0.7 mm 
lower down on the corolla tube; style 9.5 mm 
long, terete, glabrous; the two stigmatic lobes 
3.5 mm long, 2.5 mm wide, elliptic, connate % 
way up the proximal side; ovary 7 mm long, 
linear lanceoloid, glabrous, the base surrounded 
by a cupulate disk 1.8 mm high; berry (a ripe 
one) 15 mm long, 9 mm in diameter, ellipsoid, 
white, rostrate by the base of the style (one im- 
mature berry is larger, 23 X 8 mm); seeds 
0.4-0. 5 mm long, 0.23-0.25 mm in diameter, 
ellipsoid, honey-colored but the umbonate ends 
brown, the body covered with elongate poly- 
gonal cellular reticulations y 5 as long as the 
seed. 
holotypus: Hawaiian Islands, Oahu, Ka- 
hana, Waiahole Ditch Trail, Gulch no. 17, 
moist wooded gulch among Metrosideros , rare, 
800 ft alt., May 2, 1954, H. St. John 25,297 
(bish). 
discussion: C. megistocalyx is a member of 
the section Cylindrocalyces, as is its closest 
relative, C. paludosa Gaud. var. paludosa, a 
plant with the cymes 1-7 -flowered; peduncles 
3-14 mm long; pedicels 6-21 mm long; bracts 
3-8 mm long; calyx 10-20 mm long at anthesis, 
splitting into 2 lips and 5 lobes 2-10 mm long; 
corolla tube glabrous; anthers 2 mm long, the 
connective asymmetrically broad ovate; style 
6-8.5 mm long; and the stigmatic lobes 3 mm 
long. C. megistocalyx has the cymes 1 -flowered; 
peduncles 2-3 mm long; pedicels 2-6 mm long 
in flower; bracts 2.5-3 mm long; calyx 25-27 
mm long at anthesis permanently united and 
the 5 lobes represented only by minute nipple- 
like tips; corolla tube capitate glandular puberu- 
lent in the throat; anthers 3.5 mm long, the 
connective circular; style 9 mm long; and the 
stigmatic lobes 3.5 mm long. 
The new epithet is formed from the Greek 
adjective megistos, larger; and the noun calux, 
cup or calyx, and is given with reference to the 
large-sized calyx. 
