Revision of the Genus Pandanus Stickman, Part 29 
New Papuan Species in the Section Microstigma Collected by C. E. Carr 
Harold St. John 1 
The island of New Guinea has a large con- 
centration of species of Pandanus, and it is 
evident that they have been only partly collected 
and studied. They occur in nearly all plant for- 
mations from sea level to 3300 meters altitude. 
The three species here described were collected 
by C. E. Carr in 1935. He attempted to cross 
the Owen Stanley Range, from south to north, 
but lost his life in the attempt. 
Section Microstigma 
Pandanus Carrii sp. nov. (sect. Microstigma') 
Fig. 272 
diagnosis holotypi: Arboriformis 5 m alta, radici- 
bus trunco ramis syncarpioque incognitis, foliis 1.14 
(-j-0.02-0.05 ?) m longis proxima basem 3.6 cm 
latis in medio 2.6 cm latis coriaceis supra viridibus 
infra pallidioribus U-sulcatis 2-plicatis in sectione 
mediali cum 49 nervis parallelis secundariis in quoque 
dimidio nervis tertiis nullis lamina anguste gladiata 
ex basi sensim in apice crasso subulato (apex deest) 
basi inermi pallida, ex 11-13 cm marginibus cum 
aculeis 2-2.5 mm longis 6-12 mm separatis arcuatis 
fragilibus subulatis valde adscendentibus pallidis vel 
cum apicibus rubris, in sectione mediali marginibus 
cum aculeis 1.7-2 mm longis 5-12 mm separatis sub- 
arcuatis subulatis gracilibus subadpressis adscendenti- 
bus apicibus rubris, midnervo infra inermi, in apice 
subulato marginibus cum crenati-serrulis 0.2-0. 3 
mm longis 2-5 mm separatis rubris, syncarpio viridi, 
drupis 7. 7-7. 8 cm longis 1.4-1 .6 cm latis 1.1-1. 4 cm 
crassis prismaticis 6-angulosis, pileo 13-17 mm alto 
deciduo laevi exlucido dimidio infero late obtuso 6- 
anguloso dimidio supero rostrato crasso, stigmate 6-7 
mm longo late elliptico obliquo brunneo papilloso 
sulcato, endocarpio submediali 4.3 cm longo sub- 
cylindrico cuneato osseoso obscure brunneo lateribus 
lateralibus 0.7-1. 5 mm crassis intra laevibus lucidis, 
mesocarpio extra endocarpii cum fibris et eis ad stig- 
matem convergentibus, exocarpio et mesocarpio mol- 
liter carnoso et in fructu delapso et in sicco nigro, 
mesocarpio apicali extra cum fibris longitudinalibus 
sed intra cum membranis, mesocarpio basali carnoso et 
in sicco valde contracto. 
diagnosis of holotype: Tree-like, 5 m 
tall; roots, trunk, branches, and syncarp un- 
1 B. P. Bishop Museum, Honolulu, Hawaii 96819, 
U. S. A. Manuscript received March 7, 1964. 
known; leaves 1.14 (-|-0.02-0.05 ?) m long, 
3.6 cm wide near the base, 2.6 cm wide at the 
middle, coriaceous, above green, below paler 
green, U-sulcate, 2-pleated, at midsection with 
49 parallel secondary veins in each half, no 
tertiary cross veins, blade narrowly sword- 
shaped, tapering gradually from near the base 
into the stout subulate apex (the actual tip 
missing), the base pale, unarmed, beginning at 
11-13 cm up the margins with prickles 2-2.5 
mm long, 6-12 mm apart arcuate weak subu- 
late, strongly ascending, wholly pale or with 
red tips; the nearby midrib below not visible; 
at midsection the margins with prickles 1.7-2 
mm long, 5-12 mm apart, subarcuate slender 
subulate, almost appressed ascending, red- 
tipped; the midrib below unarmed; near the 
apex the margins with crenate-serrulations 0.2- 
0.3 mm long, 2-5 mm apart, red; those of the 
midrib below not observed; syncarp "green”; 
drupes 7. 7-7. 8 cm long, 1.4-1. 6 cm wide, 1.1- 
1.4 cm thick, prismatic, 6-angled; pileus 13-17 
mm high, deciduous, smooth, dull, the lower 
half forming the broadly obtuse base, this 6- 
angled, bearing a thick beak; stigma 6-7 mm 
long, broadly elliptic, oblique, brown, papillose, 
sulcate; endocarp submedian, 4.3 cm long, sub- 
cylindric, tapering downward, the apex arched, 
bony, dark brown, the lateral walls 0.7-1. 5 mm 
thick, within smooth, shiny ; mesocarp with 
thick fibers up the sides of the endocarp and 
bending in toward the stigma ; exocarp and 
lateral mesocarp soft fleshy, and at maturity 
sloughing off and drying black ; apical mesocarp 
with fibers up the sides and within partitioned 
by pale membranes and filaments; basal meso- 
carp fleshy and at maturity much shrunken. 
holotypus: Papua, Koitaki (on track to 
Owen Stanley Mountains pass), open savannah 
land, 1,500 ft alt, 13. 5. 1935, C. E. Carr 
12,274 (bm). Isotype (k) ! 
discussion : P . Carrii is a member of the sec- 
tion Micro stigma, as is its closest relative, P. 
Jiulianettii Martelli, a species with drupes 9.5- 
514 
