Page 463: Revision of Pandanus, 29 — St. John 
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loso cum seta terminali lateribus lateralibus 0.2 mm 
crassis intra laevibus lucidis, semine 9 mm longo, 
mesocarpio apicali cavernoso cylindrico cum mem- 
branis longitudinalibus brunneis paucis, mesocarpio 
basali fibroso et carnoso. 
diagnosis of holotype : Habit, roots, trunk, 
and branches unknown; leaf base and middle not 
preserved; leaf apex 39 cm long preserved, 4.5 
cm wide, thin coriaceous, above green, below 
pale green, at the point 39 cm down with 48 
parallel secondary veins in each half, the tertiary 
visible in outer part, forming narrow oblong 
meshes; blade apparently ligulate, V-sulcate, 2- 
pleated, narrowing to an acute apex, and at the 
lowest point the margins with prickles 2-4 mm 
apart, arcuate stout subulate, ascending, of two 
kinds alternating, the larger ones 0.7-1. 3 mm 
long, the smaller ones 0. 2-0.4 mm long; the 
nearby midrib below slender, elevated, and also 
with similar prickles of two types 1.5-3 mm 
apart, the larger ones 0. 2-0.4 mm long, and the 
smaller ones half as long; at the apex the 
margins and midrib below with subulate serrula- 
tions 0.1-0. 5 mm long, 0.5-1. 5 mm apart, pale; 
infructescence with a solitary syncarp; peduncle 
17 cm long, 12-15 mm in diameter, trigonous, 
leafy bracted ; syncarp completely enwrapped by 
numerous foliaceous bracts in 3 series; outer 
bracts with an expanded, lanceolate base 9-10 
cm wide, as long or longer than the syncarp, 
firm chartaceous, cream-colored, minutely prickly 
on margins and midrib below, bearing a folia- 
ceous, green, ligulate apex, probably quite long, 
but not preserved; innermost bracts 23 cm long, 
3 cm wide, slightly exceeding the syncarp, char- 
taceous, colored, linear lanceolate; syncarp 20 
cm long, 6.3 cm in diameter, cylindric, bearing 
multitudinous drupes, these 20-22 mm long, 
2.5-4 mm wide, 2.5-3 mm thick, narrowly 
prismatic, 5-6-angled, red, apparently mucilagi- 
nous throughout, upper y 7 free, ovoid, smooth, 
shiny, when dried rugose; stigma 1.5-2 mm in 
diameter, apical, flush, horizontal, subcircular, 
dark brown, papillose; endocarp in lower %, 
bony, reddish brown, tubular, with an apical 
bristle bending toward the stigma, the lateral 
walls 0.2 mm thick, within smooth, shining; 
seed 9 mm long; apical mesocarp a cylindric 
cavern with a few longitudinal brownish mem- 
branes; basal mesocarp fibrous and fleshy. 
holotypus: Papua, Koitaki, wood by stream, 
1,500 ft alt, 18. 6. 1935, C. E. Carr 12,590 
(bm). Isotype (a) ! 
discussion: P. plicatus is a member of the 
section Microstigma, as is its closest relative, 
P. rubrispicatus St. John, of northeast New 
Guinea, a species with the drupes 14-16 mm 
long, 4-6 mm wide, oblance-prismatic, the 
upper y 4 free; endocarp median; leaves 3.3 cm 
wide, and at midsection with 26 parallel secon- 
dary veins in each half. P. plicatus has the 
drupes 20-22 mm long, 2.5-4 mm wide, nar- 
rowly prismatic, the upper y 7 free; endocarp in 
lower % ; leaves 4.5 cm wide or more, and at 
the point 39 cm down with 48 parallel secon- 
dary veins in each half. 
The collector recorded, "Fruit red. Eaten by 
natives after boiling for 1 hour.” 
The new epithet is the Latin adjective plicatus, 
pleated, and is given with reference to the 
pleated leaves. 
Fig. 274. Pandanus plicatus St. John, from holotype. a, Syncarp and bracts, X 34; b, drupe, lateral view, 
X 1; c, drupe, longitudinal median section, X 1; d, drupe, apical view, X 1; e, drupe, lateral view, X 4; f, 
drupe, longitudinal median section, X 4; g, drupe, apical view, X 4; h, leaf section 39 cm from apex, lower 
side, X 1; i, leaf apex, lower side, X 1. 
