Revision of the Genus Pandanus Stickman, Part 31 
Bornean Species Collected by J. Motley 
Harold St. John 1 
Early collectors in Borneo discovered spe- 
cies of Pandanus in the native flora. Such was 
James Motley, and Martelli has already named 
P. Motleyanus in his honor. The writer has 
studied at Kew other specimens of his collect- 
ing, and has distinguished among them the 
three following new species. 
Pandanus apicalis sp. nov. (sect. Acrostigma) 
Figs. 27 6 and 276A 
diagnosis holotypi: Radices caules foliaque in- 
cogniti, pedunculo 8 cm longo 13 mm diametro 
3-laterato clavato cum syncarpio solitario, syncarpio 
6 cm diametro (licet ovoideo et 6-7 cm longo?), 
drupis 29-31 mm longis 4-7.5 mm latis 2-3 mm 
crassis fusiformibus vel oblanceoloideis acute 5-6- 
angulosis parte % supera libera et proxime curvata 
laevi vel subpunctata exlucida, stylo 3-5 mm longo 
crassiter subulato proxime curvato osseoso luteo lucido 
basi late angulosa, stigmate 3-5 mm longo lineari 
brunneo papilloso ad apicem extenso, endocarpio in 
parte 34 infera obovoideo truncato albo chartaceo 
tenuissimo, semine 4 mm longo, mesocarpio apicali 
cavernoso elongate cum fibris longitudinalibus et mem- 
branis albis, mesocarpio basali fibroso et carnoso. 
diagnosis of holotype : Vegetative parts 
unknown ; peduncle 8 cm long, 1 3 mm in diam- 
eter, 3 -sided, clavate, bearing a single syncarp, 
this 6 cm in diameter (apparently ovoid and 
6-7 cm long?) ; drupes 29-31 mm long, 4-7.5 
mm wide, 2-3 mm thick, fusiform to ob- 
lanceoloid, sharply 5-6-angled, upper y 2 free 
and proximally inclined, smooth or slightly 
punctate, dull; style 3-5 mm long, thick subu- 
late, proximally curved, bony, yellowish, shining, 
the base broadly angled; stigma 3-5 mm long, 
linear, brown, papillose, extending to the apex; 
endocarp in lower y 4 , obovoid, truncate, white, 
papery, very thin; seed 4 mm long; apical meso- 
carp an elongate cavern with longitudinal fibers 
and white membranes; basal mesocarp fibrous 
and fleshy. 
1 B. P. Bishop Museum, Honolulu, Hawaii 96819, 
U. S. A. Manuscript received March 7, 1964. 
expanded description: Plant acaulescent; 
leaves 2-3 m long, at about the middle 37 mm 
wide, near the base 28 mm wide, coriaceous, V- 
sulcate and sharply 2 -pleated, dark shiny green 
above, pale somewhat shiny green below, near 
the midsection with 34 parallel secondary veins 
in each half, the tertiary cross veins seen below 
toward the tip, forming short rhombic meshes; 
blade ligulate, tapering into the 8 cm trigonous 
subulate apex, this at the point 10 cm down 
7 mm wide, the base apparently pale and un- 
armed, beginning more than 5 cm from the base 
the margins with prickles 1.5-2. 3 mm long, 
6-14 mm apart, oblique deltoid, compressed, 
ascending, stramineous; the midrib below un- 
armed for more than the first 40 cm; near the 
midsection the margins with prickles 0.4-0. 6 
mm long, 8-13 mm apart, broad subulate to 
subarcuate lanceolate, compressed, strongly as- 
cending, stramineous; the nearby midrib below 
unarmed ; on the subulate apex the margins and 
midrib below with serrulations 0. 2-0.4 mm 
long, 1-2 mm apart, stramineous; syncarp 9-3 
cm long, 7.2 cm in diameter, ovoid; drupes 
29-38 mm long, greenish yellow, upper %—% 
free ; endocarp in lower y ; endocarp walls 
O. 1 mm thick, within smooth. 
holotypus: Borneo, grows in damp woods, 
fames Motley XXXVI, 1,247 (k mus.). 
specimens examined: Sarawak, Sabal For. 
Res., Siminjam Dist., Kuching Div., yellow 
sandy loam, lower Dipterocarp forest, 200 ft 
alt., E. F. Brunig S.9,975 (sar). 
discussion: P. apicalis is a member of the 
section Acrostigma, as is its closest relative, 
P. septentrionalis St. John, of Mount Kinabalu, 
North Borneo, a species with the drupes 26-28 
mm long; pileus 4-6 mm high, pyramidal- 
hemispheric; style 2-3 mm long; stigma 1-2 
mm long; and the endocarp apex convex. P. 
523 
