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PACIFIC SCIENCE, Vol. XXII, October 1968 
leged to examine a portion of a specimen from 
the Agardhian herbarium in Lund, which the 
Keeper, Dr. Sven Snogerup, concluded was the 
type of H. tenella, since it was annotated by 
J. G. Agardh as "Hutch tenella C. A g. sp. orig.” 
Examination of this material showed that the 
determinate branches were composed of about 
15 segments. In other respects the material 
corresponds closely with Falkenberg’s descrip- 
tion. 
H. tenella form secunda (C. Ag.) comb. nov. 
Herposiphonia secunda (C. Ag.) Falkenberg 
(1901:307) 
Hutchinsia secunda C. Agardh 1824:149 
A number of collections are assigned to this 
variously interpreted taxon, differing from typi- 
cal H. tenella chiefly in the occurrence of but 1 
determinate branch between successive indeter- 
minate branches, with 1-4 bare nodes following 
each indeterminate branch. Hawaiian islands 
— T. 934B2, 934C, on a reef near shore, Trig I., 
French Frigate Shoals, Sept. 27, 1964; John- 
ston island — T. 1084C, 25 m inside the surf 
zone, outer reef, legit R. Buggeln, Aug. 19, 
1965; line islands — L. 2708.4, tetrasporic, 
southwest reef, Jarvis I., Nov. 16, 1964; D. 
20033.1, on other algae, legit Ralph F. Palumbo, 
northeast point of Christmas I., May 16, 1962; 
TUAMOTU ARCHIPELAGO D. 11188.1, lagoon 
transect, Ngarumaoa, Raroia Atoll, legit M. S. 
Doty and Jan Newhouse, July 12, 1952; gil- 
bert islands — D. 18900.1, Abermmama I., 
legit Jane Cooper, July, 1962; marshall 
islands — G. 657.1, in the lagoon at a depth of 
35 m, Eniwetok Atoll, Sept. 7, 1955; Caroline 
islands — D. 15559-3, on Acanthophora, Koror 
I. , Palau group, legit Ernani Menez, Sept. 5, 
I960; PHILIPPINE ISLANDS — D. 18213B1, on 
Acanthophora, Mawes L, Hinatuan, Surigao, 
July 7, 1958; maldive islands — Ha. 24-21- 
132-16B, on a reef at 4 ft, Walla Faru, South 
Nilandu Atoll, Apr. 24, 1964. 
In some of the specimens listed above the 
determinate branches are somewhat larger, up to 
80p in diameter, with 40 or more segments 
1. 5-2.0 diameters long, whereas Falkenberg 
(1901) states that in H. secunda the determi- 
nate branches are relatively plump, with 14-20 
segments, which are mostly shorter than broad. 
Through the kindness of Dr. Sven Snogerup, 
Keeper of the herbarium, Lund University Mu- 
seum, the writer was privileged to examine a 
specimen from the Agardhian herbarium, which 
Dr. Snogerup considers an isotype specimen of 
H. secunda. It is a glass slide mount, no. 39158, 
with the following notation on the original 
packet: "Hutchinsia unilaterale, Aug. 6, 1825, 
Tingi, Schousboe.’’ This specimen exhibits the 
chief features described by Falkenberg (1901), 
with mostly 1 determinate branch and 2 bare 
nodes between successive indeterminate branches. 
The determinate branches are composed of 10- 
12 segments, with about 10 pericentral cells. 
The specimen is tetrasporic, with fruiting seg- 
ments about 1 diameter long. The tetrasporangia 
are in series of 7-8 in the upper part of the 
branches. The lower sterile segments are 1. 5-2.0 
diameters long. The chromatophores do not 
seem to be zonate. There are about 3 terminal 
trichoblasts, with 6-8 dichotomies. They are 
about as long as the branch bearing them. 
Fluid-preserved material from the type lo- 
cality, northern Africa, would prove helpful in 
determining the correctness of the above identi- 
fication. 
Prof. J. Feldmann of Paris kindly sent the 
writer a collection of Herposiphonia from le 
Troc, Banyuls, France. He identified this alga as 
H. secunda. The specimens were fluid-preserved 
and in excellent fruiting condition, with an 
abundance of procarps and cystocarps. In most 
respects the plants of this collection matched the 
description of H. secunda given by Falkenberg. 
However, the segments of determinate branches 
of Feldmann’ s plants are mostly 1.5— (2) di- 
ameters long, as they are in a number of the 
collections from the Pacific listed above. The 
procarps arise apically or subapically, 2-3 per 
branch, but the branch continues to grow 
beyond the developing cystocarp for 8-10 seg- 
ments. 
Falkenberg (1901) and Boergesen (1918) 
both reported instances in which the branching 
pattern for H. tenella and that for H. secunda 
occurred on the same branch. The writer has 
observed similar instances not only in plants of 
the H. tenella type, but in other species as well. 
Hence it has seemed necessary to conclude, as 
did Boergesen at first, with some reservations, 
that H. secunda should be considered merely a 
form of H. tenella. The branching pattern seems 
