Revision of Genus Alstonia — MONACHINO 
161 
usually streaked with brown; stamens inserted 
slightly above middle of corolla tube; anthers 
0.7-1. 3 mm. long. 
Type. — 'Type specimen: Mount Mariveles, 
Province of Bataan, Luzon ( 2209 Meyer), Nov. 
1904 (flower). From the same locality: ( 6876 
Elmer) Nov. 1904 (£L); ( 1164 Whit ford) 
Mar. 1905 (fr.).” 
Distribution.-— Philippines (principally in 
Luzon); a transitional form in northeast Bor- 
neo. 
Philippines. — Luzon: Borden 24037; Clem- 
ens 51889 (A); Elmer 6876 (G, NY, K); 
Loher 6512 (K), 12551 (UC), 13498 (A); 
Meyer 27198 (For. Bur. 2209; type coll.; K, 
NY, US); Ramos (Bur. Sci. 26985); Ramos 
and Edano (Bur. Sci. 48553); Whit ford 1164 
(K, NY, US). Negros: Elmer 9690 (flL; A, 
F, G, K, L, Mo, NY, L). 
British North Borneo. — Mt. Kinabalu: J. 
and M. S. Clemens 28673 (A), 28673a (A, G, 
L, UC). 
This species clearly displays affinity to A. 
angustifolia. The specimens from Borneo have 
their corolla lobes more pubescent within and 
more markedly ciliate on the margins, and are 
not streaked. They are transitional to A. an- 
gustifolia. 
18. Alstonia spectabilis R. Br., in Mem. Wern. 
Soc. 1: 76. 1811. 
A. villosa Blume, Bijdr. FI. Ned. Ind. 16: 
1038. 1826. Non Seem, FI. Vit. 161. 
1866. 
Blab er opus villosus Miq, FI. Ind. Bat. 2: 
440. 1856. 
B. villosus var. /? petiolata Miq, FI. Ind. Bat. 
2: 440. 1856. 
? Alstonia longissima F. v. Muell, Papuan 
Plants 91. 1877. 
A. villosa var. (3 glabra Koord. & Val, in 
Mededeel. Uit ’sLands PI. 11 (Bijdrage 
1): 123. 1894. 
A. somersetensis F. M. Bailey, in Queensl. 
Agric. Jour. 1: 229. 1897. 
? A. villosa forma calves cens Markgraf, in 
Bot. Jahrb. 61: 178. 1927. 
Trees up to 40 m. tall. Leaves resembling 
those of A. macrophylla, usually 3-verticillate; 
petioles 0-2 cm. long; leaf blades 10-30 cm. 
long and 2.5-9 cm. broad, varying from gla- 
brous to densely villose beneath, the lateral 
nerves 18-25 pairs, 6-12 mm. apart. Ultimate 
branches and pedicels of inflorescence usually 
ascending, not divaricate; the pedicels short, 
less than 3 mm. long; calyx lobes variable in 
size, 0.9-2. 5 mm. long, tomentose outside, 
essentially glabrous inside; corolla tube 3-3.6 
mm. long, tomentose outside; corolla lobes 
ovate to orbicular, 1.5-3 mm. long, pubescent 
outside and inside; stamens inserted at about 
l A from throat of corolla tube, the anthers 
0.9-L3 mm. long. 
Type. — "insula Timor prope Coepang, cum 
floribus fructibusque A'prili 1803 Observavi . . . 
Praecedenti [A. scholaris ] quam maxime affinis, 
sed revera distincta, nec male ab icone cit. 
Rumphii repraesentata.” Type material was not 
located at the British Museum. Although the 
interpretation of the species rests principally on 
the Timor collections cited, there is little doubt 
that Brown’s plant is here correctly placed. A. 
scholaris is also found on Timor, but Brown’s 
comments and description ("foliis quaternis 
elliptico-oblongis sub-acuminatis costatis: mar- 
gina simplicibus . . .”) definitely preclude con- 
fusion of the two. Although there is no record 
of A. macrophylla in Timor, judging by the 
distribution-range this species or a variety 
might be expected there; in any case, the in- 
florescence habit and flowers of A. macrophylla 
are so strikingly different from A. spectabilis 
and A. scholaris that there need be no fear 
of its being the original species Brown had in 
mind. The only other known species that 
conceivably might be involved is A. angustiloba, 
which is known from Java; this species is very 
similar to A. scholaris and is excluded for like 
reasons. Brown states that the illustration in 
Rumphius (1741: t. 82) is not a bad repre- 
sentation of A. spectabilis. The leaf blades pic- 
tured, their shape and acute apex, their few 
(12-15 pairs) lateral nerves, which are dis- 
