Revision of Genus Alstonia — MONACHINO 
Type of A. longissima. — "Fly-River; D’ Al- 
bertis.” Papua, 1876, No. 6424 in herbarium 
of R. Inst, studi Sup. Firenze. Described as 
glabrous, with very short petioles, leaf blades 
(ex photo of type) up to 24 cm. long and 8 
cm. broad. With a close examination of its 
pedicels and remnants of calyx lobes it may be 
possible to place this species in its proper 
position with greater confidence. 
Type of A. villosa var. glabra. — "Ex insula 
Noesabaroeng tantum nobis cognita.” 
Type of A. somersetensis. — "Near Mr. Jar- 
dine’s house, Somerset.” 
Specimens cited by Markgraf (1927) as 
A. villosa f. calves cens. — "Bismarck- Archipel: 
Neu-Pommern, ohne Fundort, fruchtend i.J. 
1901 — Parkinson 1; Gazella-Halbinsel, in den 
Waldern des Baining-Gebirges, steril Sept. 
1905 — Rechinger .3989 (Naturhis. Mus. 
Wien); Simpsonhafen, im Sekundarbusch, 
bliihend und fruchtend 16 Sept. 1908 — Ru- 
dolph 15 (einh. Name: a ituwe ): Matupi, 
bliihend Okt. 1901 — Schlechter 13684; ebenda, 
bliihend und fruchtend Sept. 1905 — Rechinger 
4114 (Naturhist. Mus. Wien). Neu-Mech- 
lenburg, Namatanai, bei Salosalo auf Schwemm- 
land, bliihend und fruchtend 15 Juni 1910 — 
Peekel 460 (einh. Name; a ituh). Kei-Inseln: 
Kei keteil a tual, mit Bliitenknospen und 
Fruchten. Aug. 1873 — leg. Beccari (Herb. 
Florenz 6413)5 I have not seen any of these 
collections. Judging from his annotated deter- 
minations of Queensland specimens and his 
published synonymy Markgraf’s concept of A. 
villosa includes A. Muelleriana. Therefore 
there is ground for doubt regarding the true 
position of this form. Markgraf noted that his 
form was not A. villosa var. glabra, as the leaves 
of the latter were said to be hardly separable 
from those of A. angustifolia, a species which 
unlike the New Guinea plant has small narrow 
leaves. As Markgraf observed, the typical leaves 
of A. spectabilis are larger than those of A. 
angustifolia, but there are frequent examples 
of small leaves in this and in all the other 
large-leaved species in the genus. Leaf size is 
not a dependable character in Alstonia. 
163 
19. Alstonia ophioxyloides F. MuelL, Fragm. 
1: 57. 1858. 
Petioles 1. 5-2.5 cm. long; leaf blades 6-12 
cm. long, pubescent beneath, the lateral nerves 
about 25 pairs, 3—6 mm. apart. Inflorescence 
and flowers like those of A. spectabilis except 
that the indumentum on the pedicels and calyces 
is of more loosely disposed and more spreading 
hairs. 
Original citation. — "Ad ripas fluviorum 
et clivos rupestres terrae Annhemicae.” 
Lectotype. — F. Mueller s. n., Fitzmaurice 
Rv. (Gray Herbarium). 
Distribution. — Australia: Northern Terri- 
tory: F. Mueller (lectotype; fl.; K, GH), F. 
Mueller (Victoria Rv.; fl.; K). 
20. Alstonia Muelleriana Domin, in Biblioth. 
Bot. 22 (Heft 89): 527. 1928. 
A. Muelleriana var. parvifolia Domin, in 
Biblioth. Bot. 22 (Heft 89): 1081. 1928. 
Trees up to 20 or 25 m. tall. Leaves usually 
3-verticillate; petioles 0-2 cm. long; leaf blades 
usually 9-18 cm. long and 3-8 cm. broad, vary- 
ing from glabrous to densely villose beneath, 
the lateral nerves 14-17 pairs, 5-13 mm. apart. 
Inflorescence with numerous crowded flowers, 
the pedicels short, ascending; calyx turbinate, 
tomentose outside, the calyx lobes ovate to 
lanceolate, densely pubescent outside and in- 
side, not reflexed; corolla tube about 2.5 mm. 
long, tomentose outside, corolla lobes linear- to 
oblong-lanceolate, about 3 or 4 times longer than 
broad, 2.2-4 mm. long; stamens inserted slightly 
above middle of tube, the anthers 0.9-1. 1 mm. 
long. 
Type. — "Endemisch in Nordost-Queensland.- 
Regenwalder bei Lake Eacham und Yarraba 
(Domin II. 1910). A. villosa F. v. Muell. 
Fragm. VI. 117 (1868). . . . Cf. descriptionem 
apud Bentham Fl. Austr. IV. 313 (1869).” 
Distribution. — Northern Australia and in 
Papua. Frequent in the northern part of 
Queensland; probably grows also in the North- 
ern Territory. Rain forests and sometimes in 
savannas, usually at low altitudes (reported up 
to 800 m. ) . 
