246 
PACIFIC SCIENCE, Vol. Ill, July, 1949 
especially at tips. No mature achenes present. 
Description of all material examined: Spread- 
ing shrub 2-6 m. tall, about 3 cm. in diameter 
at base; upper branchlets pilosulose with many 
ascending, appressed hairs. Petioles 1.0-4. 5 cm. 
long, averaging about 2 cm. long, pilosulose 
with many appressed, ascending hairs. Leaf 
blades 6-13 cm. long, 2. 5-7.0 cm. wide, averag- 
ing 9 cm. long, 4.5 cm. wide, narrow elliptic, 
elliptic-ovate, oval, or slightly obovate, thick, 
sub-coriaceous or coriaceous, very often trip- 
linerved, above glabrous, below sub-glabrous 
with only few straight, ascending, appressed hairs 
mostly restricted to veins, sometimes with very 
short, scattered hairs in intervals; margin entire, 
base cuneate and decurrent or sub-obtuse, apex 
acute or acuminate, tip of apex sharp or blunt, 
principal veins very strongly raised. Pistillate 
flowers with pedicels 0. 5-2.0 mm. long; calyx 
sub-glabrous with very few ascending, appressed 
hairs; beak attenuate apically, apex with four 
acute or acuminate teeth; stigma 2-3 mm. long, 
one side usually lacking receptive stigmatic proc- 
esses. Achene 2. 0-2. 5 mm. long, averaging 2 
mm., apical portion depressed-conic, separated 
from basal portion by abrupt obtuse constriction 
or without obvious constriction, basal portion 
smooth, thick, outer margin plane, 2. 5-3.0 mm. 
in diameter, seed ovoid. Staminate flowers with 
pedicels 0. 5-2.0 mm. long; calyx lobes thick, 
particularly at tips, pilosulose with many ascend- 
ing, appressed hairs and with or without some 
erect uncinate hairs, lobes 3-5 mm. long, 1.5- 
2.0 mm. wide, apex acute or acuminate; fila- 
ments 3. 5-5.0 mm. long, 0.5-1. 0 mm. wide, 
tapering at apex into broader, fleshy, swollen 
connective; anthers 2 mm. long, 1 mm. wide. 
Type: Gaudichaud, "In insulis Sandwicensi- 
bus, cum sequente (alt. 350-400 hex.).” (In 
Sandwich Islands, altitude 1,500-2,400 feet.) 
(Deposited in Museum National d’Histoire Nat- 
urelle de Paris [P].) 
Range: Island of Oahu, Waianae Range, moist 
regions, 2,000-3,450 feet altitude. 
Specimens examined 
Data complete: Haleauau Gulch, Mt. Kaala, 
St. John 22269 , 22210 (Ho); Mt. Kaala, Deg- 
ener 18190 (NY); Mt. Kaala, Cowan and St. 
John 333 (Ho); Puu Kalena, St. John 22368 
(Ho); Puu Kalena, Donaghho (Ho); Hapapa 
Gulch, Russ (Ho); Kanehoa, Frederick 207, 
208, 209, 210 (Ho); Kanehoa, Cowan 1034 , 
1037 (Ho); Makaha Valley, Forbes (Ho); 
Oahu, Waianae, Faurie 313 (P, Ho). 
Data not complete: Honolulu, Sandwich Is- 
lands, Hillebrand (K); Oahu, Seeman 2260 
(G, K, Ho); Mann & Brigham 220 (GH, G, 
Ho); Woahoo, Macrae (K); Gaudichaud 208 
(Bonite Voyage) (G); Remy 197, Maui (data 
very questionable) (P). 
Fig. 2. Outlines of leaves to show variation in leaf 
shape and in venation in Neraudia melastomae folia 
Gaud. var. melastomae folia Cowan (approx. y$) . 
This group seems in several ways to be inter- 
mediate between varieties uncinata and parvi- 
folia and may have arisen as a result of hybridi- 
zation between the two groups. The pedicels 
which characterize this group link it to variety 
parvifolia but the leaf shape, size, general aspect, 
and depressed-conic apex of the achene link it 
to variety uncinata. The occasional presence of 
some uncinate hairs scattered indiscriminately 
on the lower surface of the leaves further shows 
some relationship to variety uncinata. Plants of 
this group may be identified by the pedicelled 
flowers and by the thick coriaceous character of 
