25 6 
PACIFIC SCIENCE, VoL III, July, 1949 
in calyce carnoso 0.75-1.25 mm. crasso subalato 
angulatoque inclusis; semene ovato. Floribus 
masculis sessilibus vel rare pedicellatis, calycibus 
valde pilosulosis, lobis 2. 0-3. 5 mm. longis 1.0- 
1.5 mm. latis anguste navicularibus ad apicem 
longi-acuminatis, filamentis 4 mm. longis, 0.75 
mm. latis, anteris reniformibus 1.5 mm. longis 
0.75 mm. latis. 
Description of all material examined : Erect 
shrub 1. 5-3.0 m. tall, 1-4 cm. in diameter at 
base of main stem, upper branchlets with many 
erect and appressed, or only erect, hairs. Petioles 
0.8-3. 0 cm. long, with many appressed and 
erect hairs. Leaf blades 6.5-13.0 cm. long, 3.0- 
5.5 cm. wide, elliptic-ovate, ovate, or oval, thin, 
palmate or triplinerved; above sparsely sericeous, 
hairs ascending, below sericeous with more or 
less dense layer of strictly appressed or sub-erect 
hairs, oriented centripetally within primary in- 
tervals and mostly directed toward margin, 
pilosulose on raised principal veins, hairs ap- 
pressed or sub-appressed; margin entire, repand, 
partly dentate or completely dentate, base 
cuneate-decurrent to obtuse and rotund, apex 
evenly or abruptly long-acuminate or only 
acute. Pistillate flowers sessile, calyx sericeous, 
hairs appressed to erect intermixed with some 
shorter, erect, uncinate hairs, calyx beak attenu- 
ate apically with four sub-acuminate to long- 
acuminate teeth at apex; stigma 2-8 mm. long, 
without receptive stigmatic hairs on one surface. 
Achene 1. 5-2.0 mm. long, angled, apical portion 
conic, not separated from basal portion by con- 
striction, basal portion flat-convex, 1. 5-3.0 mm. 
in diameter, outer margin drying involute; seed 
ovoid. Staminate flowers sessile or with pedicels 
1 mm. or less long, calyx sericeous, hairs ascend- 
ing, lobes navicular, 2.0-3. 5 mm. long, 1.0-1.75 
mm. wide, apex long-acuminate; filaments 3-4 
mm. long, 0.3-0. 5 mm. wide; anther sacs 1.5- 
2.0 mm. long, 0. 5-1.0 mm. wide. 
This species is separated from all others by 
the sericeous condition of the plants, particularly 
of the lower leaf surface and by the conspicu- 
ously angled and ridged, fleshy calyx surround- 
ing the mature achene. The two varieties com- 
prising the species are separated on the basis of 
the posture of the hair on the lower leaf surface 
and on the leaf margin. The name for the 
species is taken from the angled character of 
the mature pistillate calyx. 
This is the group to which Hillebrand ap- 
plied Gaudichaud’s name, Neraudia sericea , 
when he reduced it to a variety of N. melasto- 
mae folia . A fragment of Hillebrand’s material, 
which upon examination proves to be what is 
named here N. angulata ; is in the herbarium of 
Bishop Museum. 
Fig. 10. Leaf outlines to show variation in leaf 
shape and shape of leaf base in Neraudia angulata 
Cowan var. angulata Cowan. 
Neraudia angulata Cowan variety angulata 
Cowan, var. nov. 
Diagnosis typi: A var. dentata differt in foliis 
marginibus semper integris subtus valde ad- 
pressi-pilosulosis sericeis. 
Description of all material examined: Shrub 
2.0-2. 5 m. tall. Petioles 1-5 cm. long. Leaf 
blades 7-13 cm. long, 3. 5-5.0 cm. wide, aver- 
aging 8 cm. long, 4 cm. wide, elliptic or elliptic- 
ovate, margin entire, base cuneate-decurrent, 
apex long-acuminate; below sericeous with more 
or less dense covering of appressed hairs, cen- 
tripetally oriented in primary intervals and 
somewhat marginally directed, producing a con- 
spicuous geometrically uneven sheen on lower 
leaf surface. 
