Genus Neraudia — COWAN 
259 
tinctness to me. He then kindly directed me to 
his locality where fourteen collections were 
made of plants of both sexes. Although the 
group had been recognized previously, the ma- 
terial available for its study was quite inadequate 
so that Mr. Degener’s assistance was a very 
definite contribution and is gratefully acknowl- 
edged. 
Neraudia kauaiensis (Hbd.) Cowan, comb. 
nov. 
Neraudia melastomae folia Gaud. var. Kauai- 
ensis Hbd., FI. Haw. Is.: 4 16, 1888. 
Upper branchlets grayish at tip with very 
many straight, erect, or sub-erect hairs. Petioles 
1-7 cm. long, with very many straight, erect 
hairs. Leaf blades 6-12 cm. long, 3. 5-7.0 cm. 
wide, elliptic, elliptic-ovate or broadly ovate, 
thin, palmate, above with few to many sub- 
appressed to erect hairs, tuft of hairs at junction 
of principal veins on upper surface erect, below 
with very many, more or less erect, irregularly 
curving and bending or almost straight hairs in 
intervals; veins densely covered with straight, 
erect hairs; margin entire or irregularly undu- 
late, base rotund or narrowing, sub-cordate, 
obtuse, or sub-obtuse, apex abruptly acuminate 
or long-acuminate. Pistillate flowers sessile, 
calyx densely pilosulose, hairs many, erect or 
sub-erect, ascending, beak attenuate apically 
with four short, acute or acuminate teeth at 
apex; stigma about 6 mm. long, receptive on all 
surfaces. Achene about 2 mm. long, apical por- 
tion conic, no constriction between apical and 
basal portions, basal portion flattened, thin, 
about 2 mm. in diameter, outer margin plane. 
Staminate flowers sessile, short-pedicelled, or 
with pedicels up to 2.5 mm. long, calyx with 
many sub-erect, ascending hairs, lobes navicular, 
3-4 mm. long, 1-2 mm. wide, with long-acumi- 
nate apex; pistil rudiment less than 1 mm. long, 
filaments flat, strap-shaped, 2. 8-4.5 mm. long, 
0.2-0. 5 mm. wide; anther sacs reniform, 0.8-2. 0 
mm. long, 0.4-1. 0 mm. wide. 
This species is easily separable from all other 
groups in the genus, except N. sericea, on the 
basis of the pubescence of the lower leaf sur- 
face. The leaves of N. kauaiensis usually have 
a sub-cordate base (except in variety Heileri 
which has irregularly undulate leaf margins) 
as compared to the usual cuneate base of N. 
sericea. The pubescence on the calices of flowers 
of both sexes in N. sericea is appressed, ascend- 
ing, whereas the pubescence on the calices in 
N. kauaiensis is erect or sub-erect, ascending. 
The venation of the leaves of the two groups 
is different and the achene of N. sericea has a 
deep, usually acute constriction whereas the 
achene of N. kauaiensis has no such constric- 
tion. 
Neraudia kauaiensis (Hbd.) Cowan variety 
kauaiensis Cowan, var. nov. 
Fig. 14 
Description of all material examined: Petioles 
1-6 cm. long, averaging 2 cm. long. Leaf blades 
6-11 cm. long, 3. 5-7.0 cm. wide, averaging 
7 cm. long, 4.5 cm. wide, elliptic-ovate or more 
often broadly ovate; above sparsely pilosulose, 
few to many sub-appressed to erect hairs pres- 
ent; margin entire, base rotund, -sub-cordate or 
obtuse. Pistillate flowers sessile. Achenes as in 
specific description. Staminate flowers sessile 
or short-pedicelled, calyx lobes 4 mm. long, 1 
mm. wide with acuminate apex; filaments 2.8- 
4.5 mm. long, 0.2-0. 5 mm. wide; anther sacs 
O. 8-2. 0 mm. long, 0.4-0. 7 mm. wide. 
Type: Rock 5323 (Ho) ( 5322-3331 [Ho, 
GH]), Kaholuamanu, Kauai. (Type deposited 
in Bishop Museum [Ho]; portions of this col- 
lection series in GH.) 
Range: Island of Kauai, wet forests. 
Specimens examined 
Data complete: On Kaholuamanu, above 
Waimea, Heller 2881 (UC, Ho, G, P, NY, 
Mich.); Kaholuamanu, Rock 17111 (Ho); 
Waimea Drainage Basin, West Side (Kokee re- 
gion — from Forbes’ field book), Forbes 1079-K 
(Ho); Kalalau Valley, Forbes 52-K (Ho). 
Data not complete: Rock 5922 (Ho); Mann 
and Brigham 624 (in part — due to the group- 
