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PACIFIC SCIENCE, Vol. Ill, October, 1949 
vel fusco-oleaginia; haustorio stuposo; stipite 
brevi, stuposo, mox in laminam latam cuneatim 
patente, terminaliter diviso in numerosa longa 
vel subdichotomopalmata segmenta cuneos latos 
simulantia; sinubus rotundatis, rursus terminali- 
ter divisis in lata linearia vel lingulata seg- 
menta; apicibus rotundatis; marginibus inter- 
gris vel perraro dentatis; sporis in series plus 
minusve longitudinales sparsis. 
Frond erect, flat, ecostate, subpalmate- 
dichotomous, not concentrically striate, up to 80 
cm. high, membranous to coriaceous, greenish - 
olive to dark olive-brown, holdfast stupose, stipe 
short, stupose, soon expanding cuneately into a 
wide blade, splitting terminally into numerous 
long, palmate or subdichotomous-palmate broad- 
ly cuneate segments, sinuses rounded, dividing 
again into terminal broadly linear or lingulate 
segments, apices rounded, margins entire or 
very sparsely toothed; sporangia scattered or in 
more or less longitudinal bands. 
Named in honour of Professor V. J. Chap- 
man, of Auckland University College, New 
Zealand, whose assistance and friendship over 
a period of many years have been greatly valued. 
The plants are probably dioecious, but the 
oogonia have not been definitely recognized. 
The antheridia are clustered into protruding, 
elongated, arched sori, up to 210 fi wide and 
60-75 [x high, containing small, cuboidal loculi. 
The sporangia, each forming a quadrant of 
spores, are large, protruding, broadly pyriform 
or clavate, with very dark content and thick 
walls, solitary or closely scattered into somewhat 
longitudinal lines, 84-108 ft high and 75-105 ft 
wide, on stalks of 1-2 cells, the latter not always 
clearly visible. Structure of from 2-3 layers of 
large, colorless medullary cells, bounded by one 
row of smaller subcortical cells and one cortical 
layer of cuboidal photosynthetic cells in width 
equal to, or half that of, the subcortical cell 
immediately below. 
There appear to be two very distinct ecologi- 
cal forms of this plant: the one greenish, mem- 
branous, broad, flabellate; the other brown, sub- 
coriaceous, narrow, dichotomous. The former 
Fig. 2. Spatoglossum chapmanii sp. nov.: a, habit 
sketch (X 0.2); b, transverse section of frond; c, an- 
theridial sorus in transverse section; d, tetrasporan- 
gium in transverse section; e, initial of tetrasporan- 
gium in transverse section; f, tetrasporangium, surface 
view. 
