18 
PACIFIC SCIENCE, Vol. IV, January, 1950 
Four others appear to require transfer from 
Myxilla into the related genus Lissodendoryx. 
They are M. behringensis Lambe, M. cratera 
Row, M. firma Lambe, and M. pygmaea 
Burton. 
Topsent (1893: xxiv) described a sponge 
from the Mediterranean coast of France as 
Myxilla arcitenens. The megascleres and 
chelas are like Myxilla, but unlike this genus 
there are not (as usual) sigmas, and there 
are (as never in Myxilla ) raphides and toxas. 
This is clearly not Myxilla. If there were no 
chelas it would fit Achiliderma. Because there 
is no genus now set up for this sponge, a new 
one is required, and is established as: 
Quintoxilla new genus 
This genus is assigned to the family Myxil- 
lidae, genotype Myxilla arcitenens Topsent. 
The holotype location is unknown to me; 
Professor Topsent told me personally that he 
rarely kept or deposited specimens. This is 
a genus with smooth dermal diactinal or 
sometimes inequiended spicules, and a prin- 
cipal skeleton of acanthostyies. The mic ro- 
se leres are isoschelas (presumably, but not 
certainly, anchorate) and toxas. Raphides 
may be among the microscleres, but this is 
not emphasized here. The name is arbitrary, 
not descriptive, suggested however by the 
emphasis upon the toxas of the spiculation. 
Naniupi new genus 
This genus is erected in the family Myxil- 
lidae, subfamily Grayellinae, with the new 
species Naniupi ula as genotype. It should 
be emphasized that this is a genus with ecto- 
somal acanthoxeas, endosomal smooth styles, 
and echinating acanthostyies. The micrc- 
scleres include arcuate chelas. The genotype 
also has peculiar sigmoid microscleres. The 
genus name is derived from the native 
Hawaiian language, "nani” meaning beauti- 
ful and "upi” meaning sponge. 
The subfamily Grayellinae is characterized 
by spiny diacts over smooth monacts. The 
type genus, Grayella, has similar ectosomal 
acanthoxeas to those of Naniupi, as very few 
sponges do, but the endosomal spicules of 
Grayella are smooth tylostyles with no echi- 
nating spicules or microscleres. Brpndsted 
( 1932: 14) described a sponge from east of 
Iceland as Grayella akraleitae. This has a 
spiculation of lumpy styles echinated by acan- 
thostyies, with arcuate chelas as microscleres. 
This cannot be a Grayella, but belongs in 
the family Microcionidae in which, however, 
a new genus is required for its reception, and 
is here established as: 
Ramosichela new genus 
This genus is assigned to the family Micro- 
cionidae, genotype Grayella akraleitae Brpnd- 
sted. The location of the holotype is not 
known to me. This is a genus for sponges 
with a principal spiculation of styles not 
smooth, in tracts that are echinated by acan- 
thostyies; there are also arcuate isochelas. 
Within this family all the other genera have 
one or more categories of smooth megascleres 
except the genus Ramoses, which has no mi- 
croscleres but is otherwise much like Ramo- 
sichela. Ramoses is exclusively Antarctic and 
sub- Antarctic whereas Ramosichela is Arctic 
or sub-Arctic. The genus name is derived 
from that of this related genus, Ramoses. 
To continue with an analysis of the sub- 
family Grayellinae, de Laubenfels (1936: 
88) puts Crellomyxilla here, but upon fur- 
ther consideration it is thought preferable to 
transfer this genus to the family Microciont- 
dae, subfamily Yvesiinae. T ethyspira has 
been put in the Grayellinae with some doubt, 
and is, still doubtfully, left there. It has small 
acanthostrongyles that may be regarded as 
dermal; its main spicules are large smooth 
styles. A fourth genus belongs in this sub- 
family, on the basis of published descriptions, 
but as it too is unnamed it is here named as 
follows: 
