Non-aquatic Frog — ATODA 
207 
found, the small and the large, seem to indi- 
cate the occurrence of sexual dimorphism in 
the species. 
SUMMARY 
1. The frog described in this article is 
commonly found in the Palau Islands. There 
are two types, large and small, apparently an 
effect of sexual dimorphism. Neither type 
possesses webs between toes or fingers. This 
frog is probably a new species of Rana. 
2. The number of eggs in one hatch is 
about 30 and they are spawned on the ground 
in weedy or bushy places. Metamorphosis 
is completed in the egg, and the frogs do not 
experience the free-swimming stage of a tad- 
pole. About 3 weeks are required for com- 
plete metamorphosis. 
3. During metamorphosis neither exter- 
nal gills nor gill clefts appear, but the ab- 
dominal skin expands to an extreme degree. 
REFERENCES 
Boulenger, G. A. 1886. Zool. Soc. Lon- 
don, Trans. 12 ( fide Brehms, Tierleben 4: 
287, 1920). 
Lynn, W. Gardner. 1940. The develop- 
ment of the skull in the non-aquatic larva 
of the tree-toad Eleutherodactylus nubicola. 
Biol. Bull. 79: 375. 
Peters, W. 1876. Akad. Wiss. Berlin, 
Monats. ( fide Brehms, Tierleben 4: 259, 
1920; Fr. Nieden, Das Tierreich 46(1): 
417-418, 1923; and H. Gadow in Cam- 
bridge Nat. Hist., 214-215, 1901). 
