Subgenera of Dubautia — St. JOHN 
341 
close relative of the other genera currently 
assigned to that subtribe, and those species 
with a smooth receptacle wholly lacking chaff 
are least satisfactorily placed there. Hence, 
the fact that all the other genera placed there 
are American does not necessarily imply that 
Dubautia is of American parentage. The 
Galinsoginae are a group transitional to the 
Helenieae. The Senecioneae also has to be 
considered, and it is worthy of note that in 
the Engler and Prantl key to the Compositae 
the old genera Dubautia and Railliardia are 
included in the keys of all three of these 
tribes. Investigations in search of a closely 
related genus have been made in these three 
tribes, but the writer must report that he 
failed to find one. 
Key to Subgenera of Dubautia 
Involucral bracts distinct; pappus bristles 
short ciliate or lacerate-ciliate . Eudubautia 
Involucral bracts united for part of their 
length, 
Pappus bristles long plumose 
Railliar diaster 
Pappus bristles short ciliate . . . Mixta 
Genus Dubautia Gaud. 
Subgenus Eudubautia subgen. nov. 
Dubautia Gaud., Voy. Freycinet Uranie, Bot. 
469, (1826) = 1830; Atlas pi 84, 1826. 
Type species: D. plantaginea Gaud., 1. c. 
Bracteae involucri liberae. Aristae pappi 
ciliati-laceratae vel breve ciliati. 
Bracts of the involucre distinct. Pappus 
bristles squamose, ciliate-lacerate, or usually 
short plumose. 
Enumeration of Species 
Dubautia Knudsenii Hbd., FI. Haw. Is. 223, 
1888. 
D. laxa H. & A., Bot. Beechey Voy. 87, 1832. 
D. laevigata Gray, Amer. Acad. Arts and Sci., 
Proc. 5: 135, 1861. 
D. magnifolia Sherff, Amer. Jour. Bot. 20: 
616 , 1933 . 
D. microcephala Skottsb., Hort. Gotoburg., 
Acta 2: 277-280, fig. 8, 1926. 
D. plantaginea Gaud., Voy. Freycinet Uranie, 
Bot. 469, (1826) = 1830; Atlas pi. 84, 
1826 . 
Subgenus Railliar diaster subgen. nov. 
Railliardia Gaud., Voy. Freycinet Uranie, 
Bot. 469, (1826) = 1830; Atlas pi. 83, 
1826. 
Type species: R. linearis Gaud., 1. c. 
Bracteae involucri cohaerentae, involucrum 
tubulosum vel anguste campanulatum infun- 
dibuliformeve est. Aristae pappi longe plu- 
mosae. 
Bracts of the involucre united into an in- 
volucre tubular or narrowly campanulate or 
funnelform. Pappus bristles usually long plu- 
mose. 
Enumeration of Species 
Dubautia arborea (Gray) Keck, Bernice P. 
Bishop Mus., Occas. Papers 11(19): 28, 
1936. 
Raillardia arborea Gray, Amer. Acad. Arts 
and Sci., Proc. 5: 134, 1861. 
D. ciliolata (DC.) Keck, Bernice P. Bishop 
Mus., Occas. Papers 11(19): 26, 1936. 
Railliar da ciliolata DC., Prodr. 6: 441, 
1837. 
D. coriacea (Sherff) Keck, Bernice P. Bishop 
Mus., Occas. Papers 11(19): 27, 1936. 
Raillardia coriacea Sherff, Bot. Gaz. 95: 
80, 1933; later as Railliardia coriacea 
Sherff, Bernice P. Bishop Mus., Bui. 135: 
122, 125, fig. 40, 1935. 
D. demissifolia (Sherff) Keck, Bernice P. 
Bishop Mus., Occas. Papers 11(19): 27, 
1936. 
Raillardia demissifolia Sherff, Bot. Gaz. 
95: 78, 1933; later as Railliardia demissi- 
folia Sherff, Bernice P. Bishop Mus., Bui. 
135: 118-120, fig. 38, 1935. 
