10 
PACIFIC SCIENCE, VoL VII, January, 1953 
of the family with respect to the three or fout 
other families. Primarily on the basis of the 
characteristics of adults of the lower genera of 
the family, Coutiere (1899) placed the cran- 
gonids below but adjacent to the Hippolyti- 
dae. Gurney (1938), on the basis of charac- 
teristics of the larvae, concluded that the 
family was most closely related to the Palae- 
monidae and of higher phylogenetic position. 
KEY TO THE HAWAIIAN GENERA 
1. Eyes at most only partially concealed by 
projection of margin of carapace, and 
visible at least anteriorly; large chela 
without a plunger on the dactylus fit- 
ting into a cavity on the finger oppo- 
site, but instead usually bearing a se- 
ries of strong teeth 2 
Eyes completely concealed by projec- 
tion of anterior carapace and visible 
only in anteroventral aspect; large 
chela almost always with a plunger 
that fits into a cavity on the. finger op- 
posite, never serrate 4 
2. Sixth abdominal segment without ar- 
ticulated pleura; large chela carried 
flexed toward body, with the palm 
excavate to accommodate merus; tel- 
son with tip emarginate 
Jousseaumea (p. 10) 
Sixth abdominal segment usually with 
pleura articulated; if large chela is car- 
ried flexed, then merus is flattened to 
accommodate palm; telson with tip 
either straight, slightly convex, or ex- 
tended to form a triangle 3 
3. Large chela carried extended; posterior 
margin of telson straight to slightly 
arcuate Alpheopsis (p. 14) 
Large chela carried flexed; posterior mar- 
gin of telson projecting as a strong 
triangle Neoalpheopsis (p. 20) 
4. With pterygostomial margin pro- 
duced into a definite angle; without 
anal tubercles; with only 2 epipod- 
ites. . . .Synalpheus (p. 26) 
With pterygostomial margin rounded, 
not angular; with anal tubercles; with 
7 to 8 epipodites Crangon (p. 46) 
Jousseaumea Weber 
Carapace extending forward as rostrum of 
variable size and as orbital teeth forming 
broad eave-like projection that conceals eyes 
in dorsal view. 
Antennular peduncle with strong stylocer- 
ite. Scaphocerite broad, oval, with squamous 
portion equal to or longer than lateral spine. 
Carpocerite of antennular peduncle short and 
broad, sometimes shorter than merocerite. 
First chela very asymmetrical, carried flexed 
at carpus so that distal end approaches body. 
Fingers of large chela serrate with large teeth 
joining exactly. Merus long. 
Carpus of second legs composed of 5 ar- 
ticles. Following legs robust with merus un- 
armed, ischium bearing 2 spines, dactylus 
simple. Sixth abdominal segment without ar- 
ticulated pleura. No anal tubercles on telson. 
Posterior margin of telson emarginate. 
Branchial formula: 5 pleurobranchs, 1 ar- 
throbranch, 8 epipodites. 
DISCUSSION: The nine species recorded for 
the world by de Man (I9II5 1922) have been 
increased to eleven by the addition of the two 
Hawaiian species described by Edmondson 
(1930). Of these, three species occur in the 
Red Sea and the Gulf of Aden, two species in 
the Banda Sea south of the Celebes Islands, 
one 'Trom the coasts of America,” one from 
Puerto Rico, and the other two species from 
the Hawaiian Islands. There are no records in- 
dicating the occurrence of any of the species 
at other than the type locality. 
HAWAIIAN SPECIES OF JOUSSEAUMEA 
1. Rostrum long and broad, extending 
from basal angles anterior to middle 
of eyes to apex beyond end of second 
article of antennular peduncle 
/. mauiensis (p. 12) 
Rostrum short, extending from base be- 
tween eyes to or slightly beyond end 
of first antennular article 
J. hrevirostris (p. 12) 
