46 
PACIFIC SCIENCE, Vol. VII, January, 1953 
tip reaching beyond end of antennular pe- 
duncle. Carpocerite dorsoventrally com- 
pressed, as long as scaphocerite. 
Large chela very large, inflated, almost cir- 
cular in section, only twice as long as broad 
at maximum diameter, with fingers much 
thinner and lower than palm, occupying dis- 
tal 0.25 of chela. Carpus of usual form. Merus 
0.25 length of chela, twice as long as broad, 
with subacute tooth terminating superior dis- 
tal margin. 
Small chela of usual configuration, with 
chela proper compressed and fingers occupy- 
ing distal 0.3. 
Carpal articles of second legs with ratio 
10 : 1.7 : 1.7 : 1.9 : 6.5. 
Ischium of third legs 0.3 length of merus; 
merus about 3.5 times as long as broad, un- 
armed; carpus 0.45 length of merus, superior 
distal margin projecting as rounded tooth, in- 
ferior distal margin not projecting but with 
slender spine; propodus 0.8 length of merus, 
somewhat curved, with 5 movable spines on 
inferior margin; dactylus 0.25 as long as me- 
rus, biunguiculate, conical inferior unguis 1.3 
times as broad at base and 0.75 as long as 
curved superior or distal unguis. 
DISCUSSION: The description and figures are 
of the type specimen previously described and 
figured by Edmondson. In this specimen, the 
only one known, the frontal regions of the 
carapace are so distorted as to render accurate 
determination of the characteristics impos- 
sible. 
This species approaches S. paraneomeris 
Coutiere in almost all characteristics. It is pos- 
sible that the rostrum and orbital hoods are 
quite different in the two species, but, as men- 
tioned, the exact condition of the frontal re- 
gion of the carapace of S. macromanus cannot 
be determined. The articles of the antennules 
may be broader in this species; the stylocerite 
may be longer; the relative sizes of the claws 
of the dactylus of the third legs may be differ- 
ent. As these characteristics are among the 
most variable in what was interpreted to be 
S. paraneomeris, I believe the range of variation 
of that species would easily encompass these 
differences. However, no specimen of S. para- 
neomeris has been seen with a large chela ap- 
proaching the inflated condition found in that 
of S. macromanus, and it appears that this is a 
valid criterion for the recognition of two spe- 
cies. Of course, it may be found that the huge 
chela of S. macromanus is merely a monstrous 
variation of 5. paraneomeris, but no specimens 
in the collection studied would indicate it; it 
may also be that the gross chela is merely a 
growth anomaly, for no other specimens have 
been collected with a similar chela. At the 
present, in any case, the species appears to be 
distinct and valid. 
DISTRIBUTION: The type specimen, a female 
13.5 mm. long, is from Lisianski Island. 
Crangon Weber 
Crangon Weber, Nomen. Ent. Sec. Ent. Syst., 
1795. 
Alpheus Fabricius, Sup. Ent. Syst., 1798. 
Carapace developed as orbital hoods which 
completely enclose eyes except on ventral 
side. Orbital hoods frequently with spiniform 
apex and usually demarked from rostral base 
by more or less pronounced depression. Ros- 
trum often present and carapace usually with 
at least slight rostral carina. Pterygostomial 
margin of carapace rounded. 
Antennules usually short, frequently with 
basal peduncular article and stylocerite re- 
duced. Scaphocerite frequently reduced; ba- 
sicerite either armed with inferolateral spioe 
or rounded; carpocerite usually reaching tn 
or beyond end of scaphocerite. 
Chela of first legs very asymmetrically de- 
veloped. Large chela of variable form, from 
subconical to compressed and twisted, sur- 
faces from smooth and entire to deeply sculp- 
tured with grooves and pronounced "alphe- 
opsidean” lobes. Dactylus usually with pis- 
ton-like process that fits into cavity on fixed 
finger and always with digital and palmar ad- 
hesive plaques. Chela always carried extended 
from body. Large cheliped with hemispherical 
