114 
PACIFIC SCIENCE, Vol. VII, January, 1953 
Second, the tooth on the inferior internal mar- 
gin of the merus of the large cheliped is re- 
duced, and the one at the corresponding loca- 
tion on the small chela is lacking in the va- 
riety, whereas both are present and well devel- 
oped in the type described by Miers. Finally, 
the first carpal article of the second legs is al- 
most twice as long as the second in C. para~ 
crinita (Coutiere: . rapport de 1.75 a 
1.85 . . loc. cit.), whereas in the variety the 
two articles are of almost equal length. 
The differences between the Hawaiian 
specimens and those described by Coutiere 
are slight except for the variations noted. 
All specimens collected in the windward 
Hawaiian Islands have been assigned to this i 
variety. It is true that the armature of the me- j 
rus of the chelipeds has been overlooked in 
making this separation, but first, the variation , 
of the armature indicates that too much faith 
should not be placed upon it; second, the Ha- 
waiian forms agree with the variety in the ratio 
of the articles of the carpus of the second legs 
— at least, they lie within the expected range 
of variation— and none had the tooth flanking 
the articulation of the dactylus of the small 
chela. If the variation in these two forms is 
similar to that found in C. brevtpes (Stimpson) 
TABLE 5 
Characteristics Differentiating the Species of the Diadema Subgroup 
CHARAC- 
TERISTIC 
C. pugnax 
C. diadema 
C. pseudopugnax 
C. percyi 
C. gracilipes 
C. paracrinita 
var. 
hengalensis 
Rostral base 
Very narrow 
triangle 
without Ca- 
rina; not 
depressed 
Broadly trian- 
gular with Ca- 
rina; some- 
what de- 
pressed 
Simple ridge 
anteriorly, 
triangular 
posteriorly; 
strongly de- 
pressed 
Slightly broad- 
er than in 
pugnax; with- 
out Carina 
anteriorly; 
strongly de- 
pressed 
Similar to 
pugnax 
Rounded, 
not well 
marked; 
slightly 
depressed 
Orbitoros- 
tral groove 
Narrow, deep 
Broad and 
shallow 
anteriorly 
Broad and 
deep 
Narrow, deep 
Narrow, deep 
Short, shal- 
low, round- 
ed 
Large chela 
Transverse 
groove* 
Transverse 
groove 
Transverse 
groove and 
depressed 
areas 
Transverse 
groove 
Transverse 
groove 
Without 
groove 
Large cheli- 
ped, merus 
Strong tooth 
on superior 
margin; in- 
ferior mar- 
gin with 
tooth and 
series of 
spines (?) 
Similar teeth, 
lacking meral 
spines 
Similar teeth, 
with serra- 
tions on all 
margins 
Same as C. 
pseudopugnax 
Similar teeth, 
2-4 spines 
on inferior 
margin 
Without su- 
perior tooth, 
inferior 
tooth weak; 
no spines 
Carpal arti- 
cles of sec- 
ond legs, 
ratio of first 
to second 
10 : 25 
10 : 10 
10 : 11 
10 : 10 
10 : 10 
10 : 10 
Third legs, 
merus 
Strong tooth 
Strong tooth 
Strong tooth 
Strong tooth 
Without 
tooth 
Without 
tooth 
*De Man states in his key ( 1911 ) "Chela without a transverse groove near the articulation of the dactylus.” However, Dana 
(loc. cit.) states "narrow emarginate above, near articulation,” and shows in his figure what might be the groove. 
