124 
PACIFIC SCIENCE, VoL VII, January, 1953 
tooth on distal inferior margin. Carpus 0.5 
as long as merus, unarmed except for tooth on 
inferior distal angle. Propodus 0.8 length of 
merus, with 10 movable spines in addition to 
usual setae. Dactylus simple, acute, somewhat 
curved. 
Telson almost twice as long as broad, tip 
slightly more than 0.5 as broad as base; lateral 
margins slightly convex anteriorly, slightly 
concave posteriorly; posterior tip strongly 
convex. Dorsal spines moderately developed, 
anterior approximately 0.4 and posterior 0.7 
of length of telson from base; medial pair of 
terminal spines narrow, acute, over 0.5 as long 
as breadth of tip of telson. Medial margin of 
tip with 3 irregularly placed, feeble spinules. 
Inner uropod with 5 spines of graduated size 
along distolateral margin; spine of outer uro- 
pod long, rather slender. 
In life, type specimen with broad, red, trans- 
verse bands. 
DISCUSSION: Only one of the paratypes had 
all its appendages; all were smaller than the 
type specimen. In these few specimens not 
much variation was noted. The most conspic- 
uous difference was in the rostral base between 
the eyes. Most of the specimens showed a 
narrow but carinate base that was set off lat- 
erally by shallow grooves, while the type 
lacked these grooves. One specimen had a 
slight spine on the merus of the small chela, 
and a specimen half the size of the type did 
not have the depressed areas on either side of 
the groove of the large chela. Aside from these 
differences and slight differences in the pro- 
portions of the articles, all the specimens were 
similar. Unfortunately there were no intact 
male specimens in the collection, but it is pre- 
sumed that the small chela of the male is ba- 
laeniceps-shaped as it is in the related species. 
RELATIONSHIP: This species plainly belongs 
to the Diadema Subgroup, and shows rela- 
tionship to C. diadema (Dana) in the nature of 
the orbital hoods and the transverse groove on 
the palm of the large chela; the rostrum is 
somewhat like that of C gracilipes (Stimpson) 
and the general configuration is sufficiently 
close to C. pugnax (Dana) that I tentatively 
identified it as that species in the field. 
C.pseudopugnax can be separated easily from 
these related species by a combination of 
characteristics. From C. diadema the species 
may be separated most easily by the nature of 
the base of the rostrum; from C gracilipes by 
the presence of a groove on the large chela 
and by the tooth on the merus of the third 
legs; from C. pugnax by the lengths of the first 
two articles of the second legs. (See Table 5.) 
This species shows a close affinity to an- 
other related Hawaiian species, C.percyi (Cou- 
tiere). They can be distinguished by the na- 
ture of the rostrum, which in C. pseudopugnax 
bears an acute carina on the anterior portion 
of the rostral base and does not overhang the 
orbitorostral groove except at the posterior 
end, while in C. percyi the anterior surface is 
broad and flat, and the margins overhang the 
orbitorostral grooves for their entire length; 
by the large chela, which bears a depression 
on the lower face in C. pseudopugnax that is 
lacking in C.percyi\ and by the small cheliped, 
which lacks teeth at the dactylar articulation 
and a tooth on the inferior internal margin of 
the merus in C. pseudopugnax^ whereas the dac- 
tylar articulation is flanked by two teeth and 
the inferior internal margin of the merus bears 
a distal tooth in C. percyi. 
This species is also related to C. philoctetes 
(de Man), but the latter has a series of mov- 
able spinules on the merus of the small chela 
and the merus of the third legs, which are ab- 
sent in C. pseudopugnax. Unfortunately the 
large chela was absent in de Man’s type speci- 
men. 
DISTRIBUTION: All Specimens known are re- 
corded above. 
Crangon percyi (Coutiere) 
Fig. 45 a-g 
Alpheus percyi Conxihs^, Soc. Philomath. Paris, 
Bui. IX, 9(5): 21, 1908. 
Alpheus percyi Coutiere, Linn. Soc. London, 
Trans. II (Zool.) 17: 426, pi. 64, figs. 22- 
