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PACIFIC SCIENCE, Vol. VII, July, 1953 
secretion (d). This secretion is the precursor 
of the veil (Fig. X’bd). As the vas deferens is 
traversed, this secretion, molded by the lateral 
contractions of the muscular wall, surrounds 
the stalks (Figs. 13^, 15c). The secretion 
from the diagonally placed epithelial cells at 
the base of the deep groove forms the viscous 
pedestal (Figs. 14^, 15c). 
The deep groove, so characteristic of the 
lumen, is no longer present in the enlarged, 
apical portion of the vas deferens (Fig. Ad). 
For a considerable distance the vas deferens 
presents a cylindrical lumen. Gradually two 
folds appear in the epithelium. From these 
typhlosole-like folds (Fig. 14c) a new secre- 
tion (/) encompasses the completed sperma- 
tophores. 
Conspicuous longitudinal muscles (Fig. 
14^) serve to move the completed sperma- 
tophores toward the genital pore. When the 
spermatophoric mass is first extruded, it is 
difficult to distinguish the pedestal from the 
veil. It is the viscous pedestal, however, which 
allows the spermatophore to become attached; 
the veil is not sticky. 
Neither the pedestal nor the veil is seg- 
mented, The spermatophore of D. asper is, 
Fig. 14, Cross section through the apical region of 
the vas deferens (Fig. Ad), a, Longitudinal muscle layer; 
b, pedestal of a single stalk; c, "typhlosole”; d, portion 
of a stalk; e, ampulla of sperm; /, mucus. (42 X.) 
a b 
Fig. 15. The completed spermatophore. a, Sheath 
of ampulla; b, spermatozoa; c, stalk; d, veil; e, pedestal. 
(35X.) 
therefore, a continuous ribbon, or two ribbons 
if both vasa deferentia expel their contents 
simultaneously. 
Comparison of Regions of Spermatophoric 
Development in D. asper with 
Those of Other Species 
Mouchet {op. cit.) assigns nine regions of 
activity to the vas deferens of Diogenes pugi- 
lator Roux. The spermatophoric differences 
observed in other species are attributed to 
vasa deferentia which lack one or more of 
these regions. The following are the nine re- 
gions of activity assigned to the vas deferens 
of D. pugilator: 
1. A rectilinear, irregular part of the canal 
coming from the testis. 
2. A right-handed helix, where the canal 
narrows, in the last turns of which the shell 
of the ampulla is secreted. 
3. A left-handed helix, where the canal in- 
creases in diameter, in the first turns of which 
the column of sperm is fragmented into seg- 
ments by its being curved into small arches 
and by ampullae being formed by the uniting 
of their extremities. 
4. In the last turns of the second helix each 
ampulla acquires a short and thick stalk. 
5. The internal canal enlarges into a spindle- 
shaped chamber. At the beginning of the spin- 
dle, in the thin part, the secretion of the 
