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PACIFIC SCIENCE, Vol. VII, October, 1953 
ADULT HAWAIIAN SQUILLIDAE 
The genera of Squillidae easily fall into two 
groups which are distinguished by differences 
in the ischio-meral articulation of the raptorial 
claw (second maxilliped) . In one group the 
articulation between the merus and ischium 
is terminal, whereas in the other the ischium 
articulates at a point anterior to the proximal 
end of the merus. This character is the basis 
for the primary division of the family, with 
Squilla, Pseudosquilla, Lysiosquilla^ and Coronida 
comprising the first group, and with Gono- 
dactylus and Odontodactylus comprising the lat- 
ter group. Although the phylogenetic sig- 
nificance of the ischio-meral articulation of 
the raptorial claw is not clearly understood. 
the species of each natural group resemble 
each other far more closely than they do those 
of the other group. 
Squilla is separated from the other members 
of the first group in that its species all possess 
longitudinal carinae on the body segments. 
Pseudosquilla, Lystosquilla, and Coronida are sep- 
arated from one another by the shape of the 
body, the carapace, and the telson, as is in- 
dicated in the following key and descriptions. 
Odontodactylus and Gonodactylus are readily dis- 
tinguished because species of Odontodactylus 
possess teeth on the inner margin of the dac- 
tylus of the raptorial claw, whereas species of 
Gonodactylus lack such teeth. 
To avoid confusion in terminology. Figure 
1 presents characteristics used in the key and 
Fig. 1. Diagrams illustrating morphological details and terminology of stomatopods. 
a, Rostrum, carapace, exposed thoracic somites, and first abdominal somite of Squilla. A-i, Structures of carapace. 
A, Rostrum; b, anterolateral spine; C, median carina; D, gastric groove; E, intermediate carina; F, lateral carina; 
G, marginal carina; H, cervical groove; I, posteriorly reflexed marginal carina. J-N, Structures of thoracic and 
abdominal somites. J, Anterolateral spine of fifth thoracic somite; k, submedian carina; L, intermediate carina; 
M, lateral carina; N, marginal carina. v-viii, Fifth to eighth thoracic somites, i, First abdominal somite. 
3, Second thoracic appendage (raptorial claw). A, Carpus; B, merus; C, propodus; D, dactylus; E, ischium. 
c, Telson and last two abdominal somites of Squilla. A-E, Structures of abdominal somites. A, 'Submedian carina; 
B, lateral carina; c, marginal carina; D, intermediate carina; E, median carina. F-L, Structures of telson. F, Pre- 
lateral denticle; G, lateral tooth; H, lateral denticle; I, intermediate tooth; j, intermediate denticles; K, sub- 
median denticles; L, submedian tooth, v-vi. Fifth and sixth abdominal somites. 
d, Last abdominal somite and telson of Odontodactylus. A-D, Structures of sixth abdominal somite (vi). A, Sub- 
median carina; B, lateral carina; c, second intermediate carina; D, first intermediate carina. E-L, Structures of 
telson. E, Marginal carina; F, second lateral carina; G, intermediate carina; H, submedian carina; I, median 
carina; J, lateral tooth; K, intermediate tooth; L, submedian tooth. 
e, Lateral aspect of head and carapace of Squilla. A, Cornea of eye; b, mandibular palp; C, antennal scale; D, first 
thoracic appendage; E, third thoracic appendage; F, second thoracic appendage (raptorial claw). 
