Revision of the Genus Pandanus Stickman, Part 14 
New Species from Malaya and Singapore 
Harold St. John 1 
As A RESULT of field work in Malaya and Sing- 
apore and of study in the herbarium at Sing- 
apore, numerous new species of Pandanus have 
been detected. The following novelties are of 
this origin. 
Pandanus attenuatus sp. nov. (sect. Acro- 
stigma) 
Fig. 150 
DIAGNOSIS HOLOTYPI: Frutex probabaliter 
humili, caule in apice 8 mm diametro lutescenti 
laevi sed cum dorsis longitudinalibus, foliis 35- 
40 cm longis 18-20.5 mm latis chartaceis supra 
viridibus infra minime viridibus pallidioribus et 
proxima basem purpureo-tinctis 1-costatis et 
minime 2-plicatis subplanis in sectione mediali 
cum 14-15 nervis parallelis secundariis promi- 
nentibus in quoque dimidio nervis tertialis sub- 
visibilis infra proxima apicem, lamina ligulata 
subito in apice caudato 5.5-6 cm longo 0.7 mm 
lato contracto basi amplexicauli inermi valde 
nervosa sed ex 2 cm marginibus cum aculeis 
0.5-1. 3 mm longis 2-6 mm distantibus subulatis 
adscendentibus pallidis, midnervo infra ultra 
mediam inermi, in sectione mediali marginibus 
cum aculeis 0.7-1 mm longis 1-4 mm distanti- 
bus subulatis valde adscendentibus, apice subu- 
lato cum marginibus et midnervo infra cum 
aculeis 0.5-1 mm longis 0.5-1 mm separatis 
subulatis adscendentibus, inflorescentia foeminea 
terminali adscendenti cum 1 syncarpio, pedun- 
culo 4.5 cm longo 4 mm diametro trigono brac- 
teato, syncarpio 3.5 cm longo 2.7 cm diametro 
ellipsoideo cum circa 144 drupis eis 13-13.5 
mm longis 4-4.5 mm latis crassisque fusiformi- 
bus 5-6-angulosis corpore 6-7 mm longo, pileo 
7.5-9 mm longo basi 3.5-4 mm alto ovoideo 
subanguloso laevi, stylo 4-6 mm longo valde 
proxime arcuato osseoso subulato olim in basi 
subanguloso, stigmate 4-4.5 mm longo late line- 
ari brunneo papilloso in apice extento, endo- 
1 B. P. Bishop Museum, Honolulu 17, Hawaii, 
U. S. A. Manuscript received June 29, 1961. 
carpio in parte l A infera lateribus cartilagineis 
pallidis, seminibus 3. 5-4.5 mm longis subdo- 
liformatis, mesocarpio apicali hemisphaerico, 
mesocarpio basali sparso cum fibris fortibus in 
lateribus. 
DESCRIPTION OF ALL SPECIMENS EXAMINED: 
Shrub, probably low; stem at apex 7—8 mm in 
diameter, yellowish, smooth but with longitudi- 
nal ridges; leaves 35-40 cm long, 18-20.5 mm 
wide, chartaceous, green above, below slightly 
paler green and near the base suffused with 
purple, 1 -ribbed, slightly 2 -pleated, but nearly 
flat, at midsection with 14-15 parallel second- 
ary veins in each half, these prominent through- 
out, cross tertiary veins barely visible only below 
and near the tip, the blade ligulate, abruptly con- 
tracted to a 5. 5-7. 5 cm caudate tip, this 0.7 mm 
wide, the base amplexicaul, unarmed, very veiny, 
beginning 2 cm up the margins with prickles 
0.5-1. 3 mm long, 2-6 mm apart, subulate, as- 
cending, pale; the midrib below unarmed to 
beyonid the middle; at midsection the margins 
with prickles 0.7-1 mm long, 1-4 mm apart, 
subulate, sharply ascending; the subulate apex 
having the margins and midrib below with 
prickles 0.5-1 mm long, 0.5-1 mm apart, sub- 
ulate, ascending; pistillate inflorescence terminal, 
ascending, bearing 1 syncarp; peduncle 3-5 — 4.5 
cm long, 4 mm in diameter, 3-sided, bracted; 
syncarp 3-3.5 cm long, 2.7 cm in diameter, ellip- 
soid, bearing about 120-144 drupes, these 13- 
13.5 mm long, 4-4.5 mm thick and wide, fusi- 
form, 5-6-angled, the body 6-7 mm long; pileus 
7.5-9 mm long, the base 3.5-4 mm high, 
ovoid, slightly angled, smooth; style 4-6 mm 
long, sharply curved proximally, bony, subulate, 
slightly angled only on the base; stigma 4-4.5 
mm long, broad linear, brown, papillose, run- 
ning to the tip; endocarp centering in lower 
but extending to the base, the walls cartilaginous, 
pale; seed 3. 5-4. 5 mm long, more or less barrel- 
shaped; apical mesocarp hemispheric, with the 
center hollow; basal mesocarp sparse, with strong 
fibers up the sides. 
3 
