Page 238: Revision of Pmdanus, 14. Malaya and Singapore — S t. John 
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smooth, gently curved, the flesh sweet, inoffen- 
sive, but not eaten; carpels 4-7, the central ones 
about 2 A as large as the marginal, the apices 
very low conic, the marginal ones asymmetric 
and some of them having just distal of the 
stigma a slight flat platform or even a concavity; 
stigmas 2.5-4 mm long, from cordate to deltoid 
or reniform, light brown, papillose, creased, 
oblique, centripetal; proximal sinus running Vi- 
2 A way to valley bottom; endocarp slightly 
supramedian, bony, mahogany-colored, the inner 
surfaces shining, the lateral walls 5 mm thick; 
seeds 12-15 mm long, 5 mm in diameter, el- 
lipsoid; upper mesocarp forming in the apex 
of each carpel a cavern with a few fibers but 
filled with aerenchyma of white medullary 
membranes; basal mesocarp fibrous and fleshy, 
extensive. 
DESCRIPTION OF STAMINATE TREE: Tree 7 
m tall, 12 cm in diameter; bark brown, muricate; 
prop roots none, but tree abused; leaves 1.23 m 
long, 3.9-4 cm wide, coriaceous, furrowed above 
the midrib, the sides not plicate, at midsection 
with 35 secondary parallel veins in each half, no 
tertiary cross veins, sword-shaped, tapering into 
a 15-20 cm subulate trigonous apex, this 10 cm 
down, 3 mm wide; at 3-4.5 cm from the base 
the margins with prickles 4-5 mm long, 7-21 
mm apart, arcuate subulate, with thickened base, 
ascending; the midrib below beginning at 6-7 
cm with prickles 3-4.5 mm long, 4-22 mm 
apart, similar but reflexed; at midsection the 
margins with prickles 1.5-2. 5 mm long, 6-12 
mm apart, subulate, closely appressed ascend- 
ing; the midrib below with prickles 1.5-2 mm 
long, 13-25 mm apart, subulate, arcuate ascend- 
ing; the subulate apex with margins and midrib 
below with prickles 0. 5-0.7 mm long, 3-7 
mm apart, subulate-serrations; staminate inflor- 
escence single, terminal, arcuate and pendent, 50 
cm long; peduncle fleshy and soon disintegrat- 
ing; lower bracts with lower part and the other 
bracts all white; lowest floral bract 48 cm long, 
2.2 cm wide, the body ligulate except at base 
with marginal prickles 2-2.5 mm long, acerose, 
ascending, closely appressed, white; the apex 
20-25 cm long, subulate, foliaceous; median 
bract 19 cm long, 2.6 cm wide, linear-lanceolate, 
unarmed except near the tip; spikes about 12, 
dense, 2.5-4 cm long, 1.5-2 cm in diameter, 
cylindric; staminate flowers very numerous, 1.5- 
1.9 cm long; column 4-8 mm long, bearing a 
common section of equal length and with about 
24 stamens, the free filament tips 1-2 mm long; 
anther body 2. 4-3. 6 mm long, linear-lanceolate, 
bearing an apical subulate prolongation of the 
connective 0.6-0. 7 mm long. 
HOLOTYPUS: Malaya, Penang Island, 5 miles 
n. of George Town, top of sea beach, June 30, 
I960, H. St. John 26,371 (bish). 
SPECIMENS EXAMINED: Malaya, Penang Is- 
land, Tanjong Tokong, 6 miles n. of George 
Town, top of sea beach, staminate, June 30, 
I960, H. St. John 26,370 (bish); Penang Island, 
Mt. Pleasure, 8 miles n. of George Town, top 
of quartz sand sea beach, clump with stems cut 
back to 1 m. by 10 cm, old phalanges from the 
ground, June 30, I960, H. St. John 26,369 
(bish). 
DISCUSSION : P. albibracteatus is a member of 
the section Pandanus, as is its closest relative, 
the Singapore species P. Boryi Gaud., which has 
the phalanges with the central apical sinuses 
1.5-2. 5 mm deep; marginal carpels with a visor 
projecting over the stigma; leaves 3. 5-4.4 cm 
wide, sword-shaped, at midsection with 39 sec- 
ondary parallel veins in each half, and near the 
base the margins with prickles 3-4.5 mm long, 
7-25 mm apart. P. albibracteatus has the phal- 
anges with the central apical sinuses 0.7-2 mm 
deep; stigmas lacking a visor; leaves 5-5.5 cm 
wide, the lower half ligulate, the outer half 
sword-shaped, secondary parallel veins 41-44 at 
midsection in each half, near the base the mar- 
gins with prickles 4.5-7 mm long and 10-45 
mm apart. 
The new epithet is formed from the Latin 
words albus, white; bracteatus, with bracts, and 
given in allusion to the white floral bracts. 
Pandanus ambiglaucus sp. nov. (sect. Pan- 
danus ) 
Figs. 156, 170 b 
DIAGNOSIS HOLOTYPI: Arbor juvenalis, 
trunco erecto 1 m alto 10 cm diametro, radici- 
bus fulturosis ad 1 m longis et 4 cm diametro 
brunneis in lineis sparse muriculatis, foliis 2- 
2.47 m longis 5.8— 6.3 cm latis coriaceis viridi- 
bus et glaucis in lateribus ambis, spinis omnibus 
albis, lamina late sulcata et dimidiis subsigmoi- 
