18 
PACIFIC SCIENCE, Vol. XVII, January 1963 
DISCUSSION: P. ambiglaucus is a member of 
the section Pandanus, as is P. inclinatus St. 
John, also of Singapore, its closest relative, and 
a species with the syncarp with 44-61 phal- 
anges, these with the apex truncate or sub- 
truncate, and the lower half fleshy enlarged; 
endocarp median; leaves 1.5— 1.7 m long, 4.4— 
4.7 cm wide, above dark olive green, below 
green; and the secondary veins at midsection 
38-40 in each half. P. ambiglaucus has the 
syncarp with 85 phalanges, these with the apex 
convex, and the lower half cuneate; endocarp in 
the lower l A\ leaves 2-2 .47 m long, 5.8-63 cm 
wide, green and glaucous on both sides; and 
the secondary veins 44-46 in each half at the 
midsection of the leaf. 
The new epithet is from the Greek words 
ambos, both; glaukos, color of the sea, applied 
to the glaucous coating on both sides of the 
leaves. 
Pandanus Boryi Gaud., Bot. Voy. La Bonite, pi. 
22, f. 15, 1841 
Fig. 157 
Description from Fosberg 36,919, Singapore: 
"Small tree,” leaves more than 1.64 m long, 
3.5 cm wide, subcoriaceous, green above and 
below, channeled above the midrib, 2 -pleated, 
in section M-shaped, at midsection with 39 
parallel secondary veins in each half, no tertiary 
cross veins, sword-shaped, tapering gradually to 
the trigonous subulate tip, more than 15 cm 
long, and about 10 cm down, only 2.5 mm wide, 
the base amplexicaul and unarmed, but begin- 
ning at 4-7 cm the margins with prickles 3- 
4.5 mm long, 7-25 mm apart, arcuate, subu- 
late, ascending, pale; the midrib below be- 
ginning at 19 cm with prickles 3 mm long, 
25-40 mm apart, similar but reflexed; at mid- 
section the margins with prickles 2-2.5 mm 
long, 8-15 mm apart, arcuate subulate, ascend- 
ing appressed; the midrib below bearing simi- 
lar prickles 1.7-2 mm long; on the subulate 
apex the margins and midrib below with ser- 
rulations 0.3-0. 5 mm long, 3-8 mm apart, red- 
tipped. . . fruiting head small cylindric, red 
when ripe”; phalange 6.3 cm long, 3.4 cm 
wide, 2.8 cm thick, pyriform, compressed, when 
dried the apex brown, but yellowish below, the 
sides smooth, shining, gently curved, free in 
upper V3, the apex low convex, lateral sutures 
none (from the direction of the basal fibers and 
from the remnant of a shoulder, it is apparent 
that when fresh the basal third had fleshy en- 
largements); central apical sinuses 1.5-2. 5 mm 
deep, broad; carpels 6, the apices very low, de- 
pressed pyramidal, the inner ones slightly the 
smaller; stigmas 2-2.5 mm wide, reniform, dark 
brown, papillose, centripetal and mostly oblique, 
some of the marginal ones with the apex trun- 
cate by an overhanging visor which terminates 
a distal concavity; proximal sinus running Vz 
way to valley bottom; endocarp median, 3 cm 
long, bony, massive, dark mahogany-colored, 
the lateral walls 3 mm thick, the inner surfaces 
shining; seeds 12 mm long, 3-4 mm in diam- 
eter, ellipsoid; upper mesocarp forming in the 
apex of each carpel a cavern with few fibers 
but filled with aerenchyma of white medullary 
membranes; basal mesocarp fibrous and fleshy. 
EXPANDED DESCRIPTION FROM OTHER SPECI- 
MENS: Branch tips 4.5-5 cm in diameter, brown, 
with crowded leaf scars; leaves 1-1.64 m long, 
3. 5- 4.5 cm wide, at midsection with 39—45 
parallel secondary veins in each half, near the 
base the marginal prickles 3-5 mm long, those 
of the midrib below 3-5 mm long; peduncle 
terminal, 22 cm long, 15 mm in diameter, 
3 -sided, leafy bracted, bearing a single syncarp, 
this 16 cm long, 14 cm in diameter, broadly el- 
lipsoid, bearing numerous phalanges; phalanges 
5.5- 6 cm long, 2-3.4 cm wide, 2-2.8 cm thick, 
pyriform to wedge-shaped, compressed; carpels 
6 - 10 . 
STANDARD specimen: Singapore: edge of 
"dry” mangrove swamp, April 17, 1956, F. R. 
Fosberg 36,919 (us). 
SPECIMENS EXAMINED: Singapore, Sungei 
Gunong, Feb. 1893, H. N. Rlidley'} 3,003 
(sing); Singapore, Dec. 15, 1904, W. Fox 
12,373 (sing). 
DISCUSSION: P. Boryi Gaud, is a member of 
the section Pandanus. The type specimen is a 
single phalange in the Paris museum, collected 
by Gaudichaud, and left without data. It was a 
misfortune that he abandoned his taxonomic 
work half done and devoted his last years to 
morphogenesis, producing lengthy papers that 
are now forgotten. P. Boryi was published with 
the binomial and with a figure showing a 
