26 
PACIFIC SCIENCE, Vol. XVII, January 1963 
14—17 mm long, 4-5.5 mm in diameter, ellip- 
soid; upper mesocarp forming in the apex of 
each carpel a cavern with a few fibers and an 
aerenchyma of white medullary membranes; 
basal mesocarp fibrous and fleshy. 
HOLOTYPUS: Singapore, Kranji Nature Re- 
serve, brackish swamp with Sonneratia caseola- 
ris, Hibiscus tiliaceus, Acrostichum aureum, 
July 10, I960, H. St. John 26,378 (bish). 
DISCUSSION : P. inclinatus is a member of the 
section Pandanus, as is its closest relative, P. 
vietnamensis St. John, of Vietnam, a species 
with the syncarp 27 cm long, with 106 phal- 
anges, these 5.8-6 cm long, the apex low con- 
vex; stigma 3.5-5 mm long or wide, black; 
leaves 0.99-1.49 m long, 5-6 cm wide, the mar- 
gin near the base with spines 4-11 mm long, 
the nearby midrib below with spines 6-8 mm 
long. P. inclinatus has the syncarp 16-23 cm 
long, with 44-6 1 phalanges, these 6-6.8 cm 
long, the apex truncate or subtruncate; stigmas 
2-3 mm long, brown; leaves 1.5-1. 7 m long, 
4.4-4.7 cm wide, the margin near the base with 
prickles 3.5-6 mm long, and the nearby midrib 
below with prickles 3-5 mm long. 
The new epithet is the Latin participle in- 
clinatus, inclined, given in reference to the ob- 
lique stigmas. 
Pandanus incrassatus sp. nov. (sect. Pandanus) 
Figs. 161, 171 
DIAGNOSIS HOLOTYPI: Arbor 7-8 m alta 8- 
10 cm diametro, cortice griseo remote muricato 
in ramis vetustioribus et trunco cum radicillis 
adventivis paucis adpresse adscendentibus, radi- 
cibus fulturosis 5-10 dm longis 10-13 mm 
diametro pallide brunneis cum radicillis brevi- 
bus spiniformatis in lineis, foliis 1.45-1.7 m 
longis 4.2-4.3 cm latis coriaceis supra olivaceo- 
viridibus infra viridibus et glaucis spinis albis 
deinde apicibus brunneis, laminis 1-sulcatis et 
cum plicis duobus rotundatis in sectione mediali 
cum 34-35 nervis secundariis in quoque medio 
nervis tertialis nullis gladiformatis sensim in 
apice 30 cm longo subulato trigono diminuen- 
tibus eo 10 cm ex apice 1.1 mm lato, basi arn- 
plexicauli et inermi albo sed ex 7—9 cm mar- 
ginibus cum spinis 3.5-5 mm longis 15-25 mm 
separatis arcuatis crassiter subulatis adscenden- 
tibus, midnervo infra ex 9-12 cm cum spinis 
3-4 mm longis 20-28 mm separatis valide sub- 
ulatis reflexis, in sectione mediali marginibus 
cum aculeis 2.5-3 mm longis 10-20 mm sepa- 
ratis arcuatis subulatis adpresse adscendentibus, 
midnervo infra cum aculeis 1.5-2 mm longis 
12-20 mm separatis arcuatis subulatis adscen- 
dentibus, in apice subulato marginibus et mid- 
nervo infra cum subulato-serrulis 0. 5-0.7 mm 
longis 3-5 mm separatis, inflorescentia foeminea 
terminali cum syncarpio solitario pendenti, 
pedunculo 23 cm longo 1.5 cm diametro trigono 
folioso-bracteato, syncarpio 15 cm longo 14.5 
cm diametro subgloboso cum 54 phalangibus 
eis 4.6-5 .2 cm longis media sup era 2.5-3. 1 cm 
longa, 3-3.6 cm lata 2.3-3 cm crassa obovoidea 
plano-truncata viridi deinde rubro-aurantiaca 5- 
6-angulosa lateribus subcurvatis infra laevibus 
et lucidis supra rugulosis et sublucidis, suturis 
lateralibus nullis, sinibus apicalibus centralibus 
plerumque (et in phalangibus lateralibus api- 
calibusque omnibus) 1-2 mm profundis late V- 
formatis rectis vel subcurvatis (vel in phalangi- 
bus subbasalibus paucis 2-3 mm profundis), 
media infera phalangis carnoso-incrassata 2.8- j 
aurantiaco-rubra infra pallidiori, humeris 2-5 
mm latis undulatis carne saccharosa sed non 
consumpta, carpelis 5-8 centralibus l /2~ 2 A tarn j, 
grandibus quam marginalibus, phalangibus di- | 
midii superi cum apicibus carpeli subtruncatis j 
vel oblatiori-pyramidalibus sed eis dimidii inferi 
cum apicibus oblato-pyramidalibus, stigmatibus j 
2-3 mm longis ellipsoideis obscure brunneis 
papillosis sulcatis obliquis prominentibus centri- 
petalibus eis marginalibus plerumque cum supra 
stigmatem galea distali cartilaginea, sinibus 
proximalibus Vi~ 2 A ad fondam extentis, endo- 
carpio 2.5 cm longo mediali osseoso obscure 
brunneo intra lucido lateribus 2.5 mm crassis, 
seminibus 12 mm longis 4 mm diametro ellip- 
soideis, mesocarpio supero in apice quique car- 
peli cavernam cum membranis albis lucidis 
medullosis formanti, mesocarpio basali fibroso 
et carnoso. 
DIAGNOSIS OF HOLOTYPE: Tree 7-8 m tall, 
8-10 cm in diameter; bark gray, remotely muri- 
cate, and the older trunk with a few appressed 
ascending adventitious rootlets; prop roots 5-10 
dm long, 10-13 mm in diameter, light brown, 
with remote, short, spinelike rootlets in vertical 
