28 
PACIFIC SCIENCE, Vol. XVII, January 1963 
rows; leaves 1.43-1.7 m long, 4.2-4.3 cm wide, 
coriaceous, above olive green, below green and 
glaucous, the spines white, becoming brown- 
tipped, 1 -ribbed and with two rounded pleats, 
at midsection with 34-35 secondary parallel 
veins in each half, no visible tertiary cross veins, 
sword-shaped, long tapering to a 30 cm subu- 
late trigonous apex, this 10 cm down 1.1 mm 
wide, the base amplexicaul and unarmed, white, 
beginning at 7-9 cm the margins with spines 
3.5- 5 mm long, 15-25 mm apart, arcuate, heavy 
subulate, ascending; the midrib below begin- 
ning at 9-12 cm with spines 3-4 mm long, 20— 
28 mm apart, stout subulate, reflexed; at mid- 
section the margins with prickles 2.5-3 mm 
long, 10-20 mm apart, arcuate subulate, ap- 
pressed ascending; the midrib below with 
prickles 1.5-2 mm long, 12-20 mm apart, arcu- 
ate subulate, ascending; near the apex the mar- 
gins and midrib below with subulate serrula- 
tions 0.5-0. 7 mm long, 3-5 mm apart; pistillate 
inflorescence terminal, bearing 1 pendent syn- 
carp; peduncle 23 cm long, 1.5 cm in diameter, 
3 -sided, leafy-bracted; syncarp 15 cm long, 14.5 
cm in diameter, subglobose, bearing 54 phal- 
anges, these 4.6-5. 2 cm long, the upper half 
2.5- 3. 1 cm long, 3-3.6 cm wide, 2.3-3 cm 
thick, obovoid, flat truncate, green, turning red- 
orange, the shade deepening downwards, 5-6- 
angled, the sides gently curving, below smooth 
and shining, above rugulose and less shiny; 
lateral sutures none; central apical sinuses 
mostly (that is, for the lateral and apical phal- 
anges) 1-2 mm deep, wide V-shaped, straight 
or gently curving (on a few subbasal ones 2-3 
mm deep); lower half of phalange fleshy en- 
larged, 2. 8-3. 6 cm wide, 2.4-3. 1 cm thick, wide 
cuneate, vermilion above, gradually paling be- 
low, the shoulders 2—5 mm wide, scalloped; 
pulp sweetish, innocuous, but not eaten; carpels 
5-8, the central 1-2 being Vl- 2 A the size of the 
marginal ones; on the phalanges of the upper 
half the carpel apices subtruncate or very oblate 
pyramidal, on those of the lower half the carpel 
apices oblate-pyramidal; stigmas 2-3 mm long, 
ellipsoid, dark brown, papillose, creased, oblique, 
prominent, centripetal, the marginal ones mostly 
with a distal, cartilaginous visor partly project- 
ing over the stigma; proximal sinus running 
Vl—Vi way to valley bottom; endocarp median 
2.5 cm long, bony, dark brown, the inner sur- 
faces shining, the lateral walls 2.5 mm thick; 
seeds 12 mm long, 4 mm in diameter, ellipsoid; 
upper mesocarp forming a cavern in each carpel 
apex with white, shining, medullary mem- 
branes; basal mesocarp fibrous and fleshy. 
holotypus: Singapore, Kranji Nature Re- 
serve, mangrove swamp, with Sonneratia caseo- 
laris, Hibiscus tiliaceus, Acanthus ilicifolius, 
near sea level, 7 July I960, H. St. John 26,374 
(BISH). 
DISCUSSION: P. incrassatus is a member of 
the section Pandanus, as is its closest relative, 
the Vietnamese species P. reversispiralis St. 
John, a species with the syncarp oval-subglo- 
bose; phalanges 4.3-4.6 cm long, orange, the 
apex low convex; stigmas all terminal, exposed; 
leaves 70-130 cm long, 4.8-6 cm wide, slightly 
glaucous above, at midsection the margins with 
prickles 3—4 mm long. P. incrassatus has the 
syncarp subglobose; phalanges 4.6-5. 2 cm long, 
red-orange, the apex truncate; marginal stigmas 
covered by a visor; leaves 145-170 cm long, 
4.2-4. 3 cm wide, olive green, not glaucous 
above, and at midsection the margins with 
prickles 2.5-3 mm long. 
The new epithet is the Latin adjective incras- 
satus, thickened, and is given with allusion to 
the thickened base of the phalange. 
Pandanus ohtusus sp. nov. (sect. Pandanus ) 
Fig. 162 
DIAGNOSIS HOLOTYPI: Arbor ? foliis 85-88 
cm longis 3.4 cm latis coriaceis 1-sulcatis lateri- 
bus arcuatis in sectione mediali cum 37-39 
nervis parallelis secundariis in quoque medio 
eis infra conspicuis, nervis tertialis nullis, lami- 
nis gladiformatis sensim in apice 15 cm longo 
subulato trigono diminuentibus eo 10 cm ex 
apice 2.5 mm lato, basi amplexicauli et inermi 
sed ex 5 cm marginibus cum aculeis 2.5-4 mm 
longis 8-14 mm separatis arcuatis subulatis pal- 
lidis in 45° adscendentibus, midnervo infra ex 
8.5 cm cum aculeis 3.5-4 mm longis 12-20 mm 
separatis subulatis valde reflexis basibus crassiter 
conicis, in sectione mediali marginibus cum 
aculeis 3-3.5 mm longis 6-11 mm separatis 
arcuatis subulatis adscendentibus apicibus ru- 
bris, midnervo infra angusto salienti cum acu- 
leis 1.5-2 mm longis 7-10 mm separatis arcu- 
