32 
PACIFIC SCIENCE, Vol. XVII, January 1963 
Pandanus rubric olor atm sp. nov. (sect. Pan- 
danus) 
Fig. 163 
DIAGNOSIS HOLOTYPI: Arboriformis 3-5 m 
alta, foliis 66-73 cm longis 2.8-3 cm latis cori- 
aceis supra viridibus infra pallidioribus supra 
midnervum sulcatis in sectione mediali cum 27 
nervis parallelis secundariis in quoque medio 
nervis tertialis nullis laminis gladiformatis in 
apice 12-15 cm iongo subulato sensim diminu- 
entibus eo 10 cm ex apice 4 mm lato basi am- 
plexicauli et inermi sed ex 3—4 cm marginibus 
cum aculeis 2.5-5 mm longis 4-18 mm sepa- 
rates subulatis pallidis adscendentibus, mid- 
nervo infra ex 8-10 cm cum aculeis 2.5-3 mm 
longis 16-30 mm separatis simulantibus sed 
adpresse reflexis, in sectione mediali marginibus 
cum aculeis 2-2.5 mm longis 4-12 mm sepa- 
ratis subulatis adscendentibus apicibus brun- 
neis, midnervo infra cum aculeis simulantibus 
1.5-2 mm longis 6-13 mm separatis; in apice 
subulato marginibus et midnervo infra cum 
subulato-serrulis 0.6-1 mm longis 2-6 mm sepa- 
ratis; inflorescentia foeminea terminal! cum 
syncarpio solitario, pedunculo 15 cm et plus 
longo 8 mm diametro exclavatis trigono folioso- 
bracteato, syncarpio 9 cm longo 8.5 cm diametro 
globoso cum circa 34 phalangibus, els 3.6 cm 
longis 2 .4-2 .7 cm latis 2-2.4 cm crassis rubris 
parte 5/9 supera libera, hac late ellipsoidea 5- 
6-angulosa apice depresse convexo lateribus sub- 
curvatis laevibus suturis lateralibus nullis, sini- 
bus apicalibus centralibus 0.5-2 mm profundis 
vadosis, carpelis 6-9 (in gemine 12 et phalangio 
3.3 cm lato) apicibus depresse pyramidalibus 
angulatis rugosis (in phalangibus basalibus 
apicibus depresse convexis) parte 4/9 inf era 
carnoso-incrassata, stigmatibus 2-3.5 mm latis 
cordatis griseis horizontalibus vel obliquis cen- 
tripetalibus, sinibus proximalibus latis Vl ad 
fondam extentis, endocarpio mediali 15 mm 
longo osseoso obscure brunneo lateribus 1.5 
mm crassis intra papillosis et sublucidis, semi- 
nibus 8-11 mm longis obliquiter ellipsoideis, 
mesocarpio supero in apice quoque carpelo 
cavernam cum fibris paucis et membranis brun- 
neis medullosis, mesocarpio basali fibroso et 
carnoso. 
diagnosis of holotype: Treelike, 3-5 m 
tall; leaves 66-73 cm long, 2.8-3 cm wide, 
coriaceous, green above, paler beneath, fur- 
rowed above the strong midrib, the sides arch- 
ing, at midsection with 27 parallel secondary 
veins in each half, no tertiary cross veins, blade 
sword-shaped, tapering upwards and gradually 
tapering to the 12-15 cm subulate apex which 
10 cm down is 4 mm wide, the base amplexi- 
caul and unarmed, but beginning 3-4 cm up 
the margins with prickles 2.5-5 mm long, 4-18 
mm apart, subulate, ascending, pale; midrib 
below beginning at 8-10 cm with prickles 2.5- 
3 mm long, 16-30 mm apart, similar but re- 
flexed, appressed; at midsection the margins 
with prickles 2-2.5 mm long, 4-12 mm apart, 
subulate, ascending, brown-tipped; the midrib 
below with similar prickles 1.5-2 mm long, 
6-13 mm apart; on the subulate tip the mar- 
gins and midrib below with subulate serrations 
0.6-1 mm long, 2-6 mm apart; pistillate in- 
florescence terminal, bearing one syncarp; pe- 
duncle more than 15 cm long, 8 mm in diam- 
eter, ' not ' clavate, 3-sided, leafy bracted; syn- 
carp 9 cm long, 8.5 cm in diameter, globose, 
bearing about 34 phalanges, these 3.6 cm long, 
2.4-27 cm wide, 2-2.4 cm thick, red, upper 
5/9 free, this part broadly ellipsoid, 5-6-angled, 
the apex low convex, the sides gently curved, 
smooth, lateral sutures none; central apical si- 
nuses 0.5-2 mm deep, shallow; carpels 6-9 (in 
a double 12, and the phalange 3.3 cm wide), 
the apices low pyramidal, angled and wrinkled 
(on the basal phalanges the carpel apices low 
convex); lower 4/9 of phalange fleshy enlarged 
as shown by persistent shoulders; stigmas 2-3.5 
mm wide, cordate, gray, horizontal or oblique, 
centripetal; proximal sinus wide, running V 2 
way to valley bottom; endocarp median, 15 mm 
long, bony, blackish brown, the lateral walls 1.5 
mm thick, the inner surfaces papillose, a little 
shiny; seeds 8-11 mm long, oblique ellipsoid; 
upper mesocarp forming in the apex of each 
carpel a cavern with a few fibers and an aeren- 
chyma tissue of brownish medullary mem- 
branes; basal mesocarp fibrous and fleshy. 
holotypus: Malaya, Pahang, Pul.au Tioman, 
Telok Paya, sea level, 19 May 1927, M. R. Hen- 
derson 18,444 (sing). 
DISCUSSION: P. rubricoloratus is a member 
of the section Pandanus , as is its closest relative, 
the Vietnamese species P. reversispiralis St. 
