Page 260: Revision of Pandanus, 14. Malaya and Singapore — S t. John 
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into a lanceolate, entire or less commonly bifid 
beak, bony, brownish, shining, flattened, proxi- 
mally curved; stigma 1 - 1.5 mm long, elliptic or 
broadly so, creased, brown, papillose, running 
almost to the apex; endocarp in lower V 3 , carti- 
laginous, brown, walls 0.1 mm thick, the inner 
surface pale, dull; seed 3-4 mm long, ellipsoid, 
! truncate; apical mesocarp a slender cavity 9 mm 
long; basal mesocarp fibrous up the sides, fleshy 
within, 4-5 mm long. 
HOLOTYPUS: Malaya, Johore, Sungei Kayu, 
in swamp, 17 Oct. 1936, Kiah bin Haji Salleh 
32,102 (SING). 
SPECIMENS EXAMINED: Malaya, Sfungei] 
Kayu, Mawai-Jemaluang Road, in swamp, evi- 
dently fruits commonly, 11 Oct. 1936, E. ]. H. 
Corner 32,460 (SING), in large part, but not 
the loose fruit), (sing). 
DISCUSSION : P. brevicornutus is a member of 
the section Rykia, as is its closest relative, the 
Malayan species P. kedahensis St. John, which 
has the syncarp 12.5 cm long, 11 cm in diam- 
eter; drupes 4. 2-4.4 cm long, 8-13 mm wide, 
narrowly oblanceoloid; style 4-5 mm long, 
bifurcate, the lobes mostly divergent; endocarp 
j with apical narrow pyramidal projection; seeds 
15- 16 mm long; leaves 4.5-4.6 cm wide, and 
at midsection with 43 secondary parallel veins 
in each half. P. brevicornutus has the syncarp 
3 . 5 - 5.2 cm long, 3-4 cm in diameter; drupes 
16- 18.5 mm long, 2 - 2.5 mm wide, narrowly 
1 cylindric; style 3-4 mm long, lanceolate, less 
commonly unevenly bifid; endocarp apex trun- 
cate; seeds 3-4 mm long; leaves 24-29 mm 
wide, and at midsection with 18-21 secondary 
parallel veins in each half. 
The new epithet is taken from the Latin 
words brevis, short; cornutus, horned, and is 
chosen in reference to the short, hornlike style 
of this species. 
Pandanus distentus sp. nov. (sect. Rykia) 
Fig. 165 
DIAGNOSIS HOLOTYPI: Planta cum foliis 1.8- 
3 m longis 9 cm latis crassiter coriaceis 1 -sul- 
; catis 2-plicatis in sectione M-formatis in sec- 
tione mediali cum 61 nervis parallelis secunda- 
riis prominentibus in quoque medio nervis ter- 
1 tialis fortibus in dimidia ultima reticulam for- 
mantibus lamina ligulata valde contracta in 
apice 29 cm longo subulate caudato trigono eo 
10 cm ex apice 1 mm lato basi non conservati, 
proxima basem marginibus cum spinis 5-7 mm 
longis 11-24 mm separatis deltoideo-subulatis 
complanatis subarcuato-adscendentibus pallidis 
sed apicibus nigris, midnervo infra cum spinis 
teretihus 5-6 mm longis 12-30 mm distantibus 
subulatis arcuatis reflexis nigris basibus crassis, 
in sectione mediali marginibus cum spinis 4- 
5.5 mm longis crasse subulatis arcuatis adscen- 
dentibus saepe tota pallidis, midnervo infra cum 
spinis 4 mm longis 4.5-6 cm separatis crasse 
subulatis arcuatis complanatis valde reflexis, in 
sectione diminuenti marginibus cum subulato- 
serris 1 . 2-2 mm longis 1-4 mm separatis arcu- 
atis, midnervo infra angusto salienti cum aculeis 
simulantibus, in apice caudato marginibus et 
midnervo infra cum serrulis 0.2-0.4 mm longis 
1-4 mm separatis, inflorescentia foeminea "circa 
30 cm longa,” pedunculo 10 mm diametro 
trigono folioso-bracteato probabaliter cum syn- 
carpio solitario, syncarpio circa 15 cm longo et 
10-11 cm diametro ellipsoideo, drupis multis 
34-36 mm longis (vel stylo sequi 37-41 mm) 
8-12 mm latis 7-12 mm crassis anguste oblan- 
ceoloideis subcompressis 5 - 7 -angulosis corpore 
30-32 mm longo anguste ellipsoideo vel fili- 
form! lateribus subcurvatis vel planis, pileo 9 - 
12 mm longo basi pyramidali-semiorbiculari 
laevi, stylo 3-5 mm longo osseoso rigido lucido 
compresso proxime curvato basi late oblonga 
parte supera acute bifida cum lobis subulatis 
sed eis cum lateribus exterioribus extensis 
undulatis, stigmate 3-4 mm longo late ovoideo 
bifido ad apices extenso brunneo papilloso, 
endocarpio submediali osseoso obscure brunneo 
lateribus 1 mm crassis intus lucido in apice cum 
projectione central! anguste conico, semine 15 
mm longo 5 mm diametro ellipsoideo obtuso, 
mesocarpio apicali cavernoso cum fibris et 
membranis, mesocarpio basali in lateribus fibro- 
so intra carnoso. 
diagnosis OF HOLOTYPE: Stature not re- 
corded; leaves 1 . 8-3 m long, 9 cm broad, thick 
coriaceous, 1 -ribbed and 1 -furrowed, 2 -pleated, 
in section M-shaped, at midsection with 61 
parallel secondary veins in each half, these rather 
prominent except at base, the tertiary cross 
veins strong, transverse, forming oblong meshes, 
these at least from the middle outwards giving 
