Page 264: Revision of Pmdanus, 14. Malaya and Singapore — St. John 
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3-planato cum circa 832 drupis eis 17—19 mm 
longis 2-2.5 mm latis 1.5-2 mm crassis viridi- 
bus corpore 14-15 mm longo anguste cylindrico 
compresso 5-6-anguloso, pileo 6-8 mm longo 
basi 2-2.5 mm alta ellipsoidea laevi, stylo 5-6 
mm longo plerumque proximo frequente subre- 
flexo, stigmate 2-3.5 mm longo anguste ellip- 
tico brunneo papillose ex apice distanto, endo- 
carpio in parte l A infera cartilagineo pallide 
brunneo lateribus 0.1 mm crasso intra exlucido, 
semine 2.7 mm longo ellipsoideo, mesocarpio 
apicali cavernoso 8 mm longo, mesocarpio basali 
in lateribus fibroso intra carnoso. 
DESCRIPTION OF ALL SPECIMENS EXAMINED: 
Shrub up to 1.7 m tall, 8-15 mm in diameter, 
often decumbent; bark dark mahogany-colored, 
shining; internodes 7-11 mm long; prop roots 
several and as much as 40 cm long, 7-8 mm in 
diameter, brown, with loose spirals of adventi- 
tious rootlets 2-5 mm long, divergent, prickle- 
like; leaves 37-65 cm long, 25-34 mm wide at 
the middle, 20-24 mm wide near the base, 
chartaceous, above glossy dark green, below yel- 
lowish green, at base channeled above the mid- 
rib, slightly 2 -pleated, ligulate, the apex nar- 
rowed to a 4-5 cm trigonous subulate tip 1-1.5 
mm wide, at midsection with 18-22 secondary 
parallel conspicuous veins in each half, but no 
clearly visible tertiary veins, the base amplexi- 
caul, unarmed, but beginning at 3-3.5 cm the 
margins with prickles 2. 5-3. 5 mm long, 4-8 mm 
apart, subulate, bent upwards, brownish; the 
midrib below beginning at 10 cm with a few 
remote similar prickles 3 mm long, reflexed; at 
midsection the margins with prickles 0.8-1 mm 
long, 7-20 mm apart, subulate, appressed as- 
cending; the midrib below with one or several 
remote prickles 1.5 mm long, compressed, heavy 
based, arcuate ascending; on the subulate tip the 
margins and midrib beneath with prickles 0.3- 
0.5 mm long, 0.5-1. 5 mm apart, subulate, as- 
cending; infructescence terminal, with a single 
head, or rarely two heads; peduncle 10-17 cm 
long, 4-6 mm in diameter, trigonous, leafy 
bracted, sometimes pendent; syncarp 4-6 cm 
long, 3-4.5 cm in diameter, ellipsoid but slightly 
3-sided, bearing 832-1,040 drupes, these 17-19 
mm long, 2-2.5 mm wide, 1.5-2 mm thick, 
green, the body 14-15 mm long, narrowly cylin- 
dric, compressed, 5-6-angled; pileus 6-8 mm 
long, the base 2-2.5 mm high, ellipsoid, smooth; 
style 5-6 mm long, mostly proximal in attach- 
ment, often somewhat reflexed; stigma 2-3.5 
mm long, narrowly elliptic, brown, papillose, 
remote from the apex; endocarp in lower 
cartilaginous, pale brown, the walls 0.1 mm 
thick, the inner surface dull; seed 2.7 mm 
long, ellipsoid; apical mesocarp a cavern 8 mm 
long; basal mesocarp fibrous up the sides, fleshy 
within. 
HOLOTYPUS: Malaya, Johore, Sungei Sedili, 
30 July 1939, Kiah bin Haji Salleh 63,926 
(SING). 
SPECIMENS EXAMINED: Malaya, Johore, 5Vl 
miles, Kota Tinggi-Mawai Road, in very swampy 
places in backwaters of streams, forming more 
or less pure stands, 13 April 1935, E. J. H. 
Corner 29,231 (BO, sing); Johore Sfungei] 
Berassau, Mawai-Jemaluang Road, frequent in 
swampy forest, 6 Feb. 1935, E. J. H. Corner 
28,744 (sing); ditto, flopping, gregarious in wet 
hollows in swampy forest, 7 Feb. 1935, Corner 
s. n. (SING); ditto, gregarious in standing water, 
low elevation, 28 April 1935, Corner 29,361 
(BO, sing); ditto, in standing swamp by the 
stream, 5 Jan. 1936, Corner 29,991 (sing). 
discussion: P. inundatus is a member of the 
section Rykia and in that section is its closest 
relative, P. Scortechinii Martelli, a species with 
drupes 12-13 mm long; style terminal, 4-5 mm 
long; stigma linear; endocarp slightly subme- 
dian; seed 6 mm long; leaves 25-27 cm long, 
near the base 14 mm wide, at midsection the 
secondary nerves 15-16 in each half, tertiary 
cross veins visible in the outer half, at midsec- 
tion the margins with weak prickles 0.2-0. 3 
mm long, flat appressed, close or remote or none. 
P. inundatus has the drupes 17-19 mm long; 
style of lateral drupes markedly lateral, 5-6 mm 
long; stigma narrowly elliptic; endocarp in lower 
Vy seed 2.7 mm long; leaves 37-65 cm long, 
near the base 20 mm wide, at midsection the 
secondary nerves 18-22 in each half, no clearly 
visible cross veins; at midsection the margin 
with prickles 0.8-1 mm long, 7-20 mm apart, 
subulate, appressed ascending. 
This was probably the species illustrated by 
Martelli (Webbia 4(2): t. 32, f. l-3a, 1914) 
as his P. johorensis Martelli. Martelli did not 
have the habit of indicating the specimen from 
