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PACIFIC SCIENCE, Vol. XVII, July 1963 
Papy was aware of this. Neither Moore nor 
Grant found F. nutans on Raiatea, and Papy 
was aware of the description of F. cuneata from 
that island. Moorea is certainly a likely locality 
for F. nutans, but for none of these localities 
does Papy cite specimens. 
Thus, the geographic distribution offered by 
Carlquist (1957:2) must be broadened to in- 
clude the occurrence of F. nutans on Tahiti-iti, 
F. cordata on Bora Bora, and F. cuneata ssp. 
tahaaensis on Tahaa. 
4. Fitchia rapensis F. Brown, Bishop Mus. Bull. 
130:366. 1935. 
Since the ten previously recorded collections 
of this species are all without flowers (Carl- 
quist, 1957 : 62 ) , the description of a flowering 
specimen should be of interest: Rapa, moist 
zone, altitude 520 m, Jan. 3, 1922, W. B. Jones 
372 ( Whitney Expedition) ( BKL ) . 
"Tree,” glabrous. Petioles 40-75 mm long. 
Blades ovate, the larger ones averaging 90 mm 
long and 84 mm wide, rounded or scarcely 
cuneate at the base, the apex acute; entire; with 
about 12 pairs of major lateral veins. The sin- 
gle inflorescence is terminal, with two heads, 
the peduncles 50 and 60 mm long, reflexed. 
Heads 45 mm long and 45 mm wide. Bracts in 
about 4 series, those of the second layer semi- 
circular, 10 mm long and 22 mm wide, and 
those of the inner layer 20 mm long and 12 
mm wide. Receptacular bracts cuneate, 16-17 
mm long, 3-4 mm wide. Mature corollas 24-26 
mm long, the ventral cleft reaching to 6-8 mm 
of the base, the segments eventually separating 
to within 10 mm of the base; the tips of the 
lobes often densely covered with sclerified cells, 
which are up to 0.75 mm long. Anther sacs 
5.5-6 mm long, caudate, the tail 0.3-0.4 mm 
long; the stamen tips cuneate, 1.5 mm long. 
Mature style ( broken off) at least 32 mm 
long. Achenes, at and just after anthesis, 10-12 
mm long, 4 mm wide, the awns 12-15 mm 
long, strongly bent and twisted at the base. 
ANATOMICAL DESCRIPTIONS 
Anatomical studies were based on the her- 
barium specimens cited above. Liquid-preserved 
heads of F. cuneata ssp. tahaaensis and wood 
samples of both F. cuneata ssp. tahaaensis and 
F. cordata , for which the herbarium specimens 
serve as vouchers, were available. The methods 
of preparation of these materials are the same 
as those employed for such materials in the 
two papers (Carlquist, 1957, 1958) dealing 
with the anatomy of this genus. 
SECONDARY XYLEM: With respect to quali- 
tative features, the species of Fitchia other than 
F. speciosa are much alike in wood anatomy 
(Carlquist, 1958). Fitchia cuneata ssp. tahaaen- 
sis and F, cordata share the qualitative features 
of those species. The bands of apo tracheal 
parenchyma are clearly shown by the two new 
taxa ( Figs. 4, 5 ) . Thick-walled fibers are readily 
apparent In F. cuneata ssp. tahaaensis (Figs. 1, 
3 ) ; libriform fibers are less thick-walled in 
F cordata (Figs. 5, 6). Sclerosed tyloses have 
been reported in F. nutans and F. speciosa 
(Carlquist, 1957, 1958); they are abundant in 
F. cuneata ssp. tahaaensis (Figs. 1, 2), although 
none were observed in F. cordata . 
Quantitative features of the two new taxa 
are as follows: F. cordata : vessels per group: 
1.37; average vessel diameter: 61.5 /x; diameter 
of widest vessel: 88" "p; average length vessel 
elements: 325 ft; average length libriform fibers: 
502 /a; average length apotracheal parenchyma 
cells: 356 g; average height multiseriate rays: 
1.17 mm. Fitchia cuneata ssp. tahaaensis: ves- 
sels per group: 1.66; average vessel diameter: 
62.8 /a; diameter of widest vessel: 92 /a; average 
length vessel elements: 326 /a; average length 
libriform fibers: 507 /a; average length apo- 
tracheal parenchyma cells: 380 /a; average 
height multiseriate rays: 1.51 mm. 
Quantitative as well as qualitative data point 
up a close similarity between F. cuneata ssp. 
tahaaensis and the typical F. cuneata. Quanti- 
tative and qualitative data for F cordata re- 
FIGS. 17-19. Fitchia cuneata ssp. tahaaensis. Fig. 17: Transection of involucral bract taken about midway 
along length; adaxial face below. Fig. 18: Transection of receptacular bract; ad axial face and adjacent achene at 
left. Fig. 19: Transection of style. At center of style, stigmatoid tissue. Exterior to each bundle is a secretory 
canal, except for bundle at upper left, which is flanked by a pair of canals. At right, anthers in transection. 
Note secretory canal in each of the two anthers. Fig. 17, X 100; Fig- 18, X 80; Fig. 19, X 120. 
