Page 274: Revision of Pandanus, 15. Malaya — St. John 
331 
cm with prickles 3-3.5 mm long, 8-20 mm 
apart, very stout subulate, strongly reflexed; pis- 
tillate scape 15-40 cm long, bearing 3-9 syn- 
carps, these 5-7 cm long; drupes very hard until 
ripe when they become bright reddish orange 
and soft. 
LECTOTYPE: Singapore, Bukit Timah [Bukit 
Mandai Road], 1894, H. N. R[idley ], (SING). 
Specimen examined and herein designated as 
lectotype. 
SPECIMENS EXAMINED: Singapore, Pasir Pan- 
jang, 17 Oct. 1912, I. H. Bur kill & M. R. Hen- 
derson 6,802 (bo, SING). Malaya: Johore, Pulau 
Kukub, 1904, H. N. Ridley (paratype), (SING). 
Ridley, when publishing this species, cited 
three collections and mentioned Sarawak, but 
did not designate a holotype. The most com- 
plete of his original specimens is here chosen 
as the lectotype. 
Pandanus collinus Ridl., Mater. FI. Malay Penin. 
2: 228, 1907; FI. Malay Penin. 5: 79, 
1925; Martelli, Webbia 4(1): 10, 1913; 
(sect. Acrostigma ) 
Fig. 173 
DIAGNOSIS OF LECTOTYPE: Shrub 1. 2-2.1 m 
tall; stem at summit 6 mm in diameter, brown- 
ish, smooth; leaves 25-45 cm long, 8-11 mm 
wide, firm chartaceous, apparently pale green 
on both sides and flat except near the base 
where 2 -pleated, 1 -ribbed, above unarmed, at 
the middle with 10-11 secondary parallel veins 
in each half, below the tertiary cross veins 
barely visible, gradually narrowed in the last 
10 cm to a subulate tip 15-20 mm long and 
0.6-0.8 mm wide, the base amplexicaul, un- 
armed, blackish purple, beginning at 2 cm 
from the base the margins with prickles 0.7-1 
mm long, 3-7 mm apart, subulate, ascending, 
yellowish but red-tipped; the midrib below un- 
armed to well beyond the middle; at mid- 
section the margins with subulate tipped ser- 
rulations 0. 2-0.3 mm long, 1.5-3 mm apart, 
appressed ascending, red-tipped; near the apex 
the margins with serrulations 0. 1-0.2 mm long, 
0.3-0. 5 mm apart; the midrib below with ser- 
rulations 0. 2-0.3 mm long, 0.5-1. 5 mm apart; 
pistillate inflorescence terminal, erect, bearing 
1 head; peduncle 8 cm long, 3 mm in diameter, 
bracted; syncarp 3 cm long, 2.5 cm in diameter, 
subglobose, bearing about 88 drupes, these 
14-15 mm long, 4-5 mm wide, 3-4 mm thick, 
the body ellipsoid to obovoid, 5-6-angled, 10 
mm long; pileus with the base 3.5-4 mm high, 
semiorbicular or slightly narrower, rather 
smooth, but with longitudinal ridges and lines; 
style 4-5 mm long, abruptly subulate, angled, 
sharply proximally curved, bony; stigma 3-4 
mm long, elliptic-linear, brown, papillose, ex- 
tending to the tip; endocarp centering in lower 
Vb but extending to the base, the walls 0.1 mm 
thick, cartilaginous, stramineous; seed 7.5-8 mm 
long, lance-obovoid; upper mesocarp an oblate 
hemisphere, filled with white medullary mem- 
branes; basal mesocarp fibrous up the sides, 
fleshy within, sparse. 
LECTOTYPE: Malaya, Kedah, Kedah Peak, 
woods, 1893, H. N. R[idley }, (SING). Specimen 
examined and here designated as lectotype! 
DISCUSSION: P. collinus belongs in the section 
Acrostigma. It was described by Ridley from 
three collections from Malaya. The best of these 
and the one labeled P. collinus! by Ridley is 
here chosen as the lectotype. Contrasting dif- 
ferences are given for this in our treatment of 
P. alticola. 
Pandanus glaucophyllus Ridl., Roy. Asiat. Soc., 
Straits Branch, Jour. 41: 50, (1904) = 
1903; ( sect. A cro stigma ) 
Fig. 174 
DIAGNOSIS OF holotype: "Stems 3-4 m. tall, 
12-25 mm. in diameter,” the internodes brown 
to yellowish, smooth, mostly higher than the 
nodes; leaves 52-60 "or more” cm long, near 
the base about 4 cm wide, near the apex 5. 7-6.5 
cm wide, appearing narrowly oblong, rather 
flaccid and when dry chartaceous, glaucous, 1- 
ribbed but 2 -pleated and in section low M- 
shaped, the secondary parallel veins conspicuous 
and 34 of them in each half, the apex abruptly 
contracted to a 6-7 cm trigonus subulate caudate 
prolongation which near its base is 1 mm wide, 
the base of the blade amplexicaul and unarmed, 
but beginning at 4 cm up the margins with 
prickles 0. 5-0.8 mm long, 1-3 mm apart, slen- 
der subulate, pale, ascending or diverging and 
some of them doubles or triples; the midrib 
