Page 280: Revision of Pandanus, 15. Malaya — S t. John 
337 
with prickles 0.5-1 mm long, 0.3-1 mm apart, 
subulate, diverging or subascending; the mid- 
rib with stouter prickles 0.7-1 mm long, 0.5-1 
mm apart, slightly ascending; on the caudate 
tip the margins and midrib below with prickles 
0.5-0.9 mm long, 0.5-1 mm apart, arcuate 
subulate, ascending; median bracts 3-4.5 cm 
long, 1.6-2 A cm wide, lanceolate, the margins 
and the midrib below towards apex minutely 
prickly; spikes 5-8 and 1.5-2. 5 cm long, 0.8- 1.3 
cm in diameter, ellipsoid, dense; stamens very 
numerous, attached singly and directly to the 
axis; filament 0.2 mm long; anther 4.5-5 mm 
long, 0.2 mm wide, bearing an apical subulate 
prolongation of the connective 0.4-0.7 mm 
long. 
HOLOTYPE: Singapore, Kranji, 1892, H. N. 
R[idley J, (SING). Holotype examined! 
SPECIMENS EXAMINED: Singapore, Hinterin- 
dien, Dec. 16, 1905, A. Engler 3,862 (b); Singa- 
pore, Bidadari, 1898, H. N. R{idley ] 9, 171 
( FI ) ; Singapore, cult., first male flowers seen, 
1895, H. N. R[idley ], (fi, sing); without col- 
lector, ex Hb. (SING), Forest Iurong, low grow- 
ing plant, Jan. 10, 1889, Flora of Singapore 92 
( B ) , the holotype of P. flagellifer. 
Malaya: Johore, Sungei Pelepah Kiri, 26 June 
1928, E. J. H. Corner 33,336 (sing). 
Pandanus perakensis Ridl., Mat. FI. Malay Penin. 
2: 231, 1907; FI. Malay Penin. 5: 81, 
1925; (sect. Aero stigma) 
Fig. 177 
diagnosis of LECTOTYPE: (Probably acau- 
lescent); leaves 1.5-1. 8 m long, 2-3.6 cm wide, 
subcoriaceous, light green, linear, rather ab- 
ruptly narrowed to the 9 cm subulate, trigonous 
apex, this 10 cm down 6 mm wide, in section 
the blade AA-shaped, at midsection with 39—41 
secondary veins in each half; near the base the 
margins with spines 2-3 mm long, 5-13 mm 
apart, stout subulate, arcuate, pale, broad based, 
ascending; the nearby midrib below with spines 
2.5-3 mm long, 6-13 mm apart, similar but 
even thicker subulate and reflexed; at mid- 
section the margins with subulate-tipped serrae 
0.5-1. 5 mm long, 4-8 mm apart, the midrib 
below with prickles 0.5-1 mm long, 8-15 mm 
apart, subulate; near the apex the margins with 
serrulations 0. 3-0.5 mm long, 2-3 mm apart; 
the midrib below with similar ones but 2-4 
mm apart; peduncle 15 cm long, 8-9 mm in 
diameter, straight, bracted; syncarps 3-4, spicate, 
crowded, 4. 5-6. 5 cm long, 3. 5-4. 5 cm in di- 
ameter, elliptic-cylindric (rarely subglobose), 
appearing spiny from the numerous curved as- 
cending styles ( but a few at the very base 
reflexed), bearing about 430 drupes; drupes 
20-22 mm long, 5-6 mm wide, 4-5 mm thick, 
oblance-fusiform, compressed, apparently green, 
upper l /z free, the body 13-14 mm long, ob- 
lanceoloid, the apex truncate, pileus at last 
shedding, the base semiorbicular, bearing a style 
6-7 mm long, stout subulate, ridge angled, 
gently arcuate towards apex of syncarp; stigma 
4-5 mm long, linear, distal, running to the very 
apex; endocarp 6-7 mm long, centering in 
lower V3, ellipsoid, truncate, stramineous, car- 
tilaginous, the sides 0.15 mm thick; apical meso- 
carp 3-3.5 mm long, a fat discoid cavern with 
a few white membranes; basal mesocarp fibrous 
along the margins, fleshy within. 
EXPANDED DESCRIPTION: Pistillate plants 
with peduncle 13-15 cm long; syncarps 2-3 
and 4.5-7 cm long, 3. 5-5. 5 cm in diameter. 
LECTOTYPE: Malaya, "Perak, Maxwell’s Hill”; 
June 1893, H. N. R{idley ], (sing). Lectotype 
examined! 
SPECIMENS EXAMINED: Malaya, Pahang, Ku- 
ala Lipis, 1892, A. S. Machado 11,633 (SING); 
Kedah, Baling Hill, 21 Nov. 1941, E. J. H. 
Corner & J . C. Nauen (SING). 
Pandanus spinulosus (Ridl.) comb. nov. (sect. 
Acrostigma) \ P. collinus Ridl. var. spinu- 
losus Ridl., FI. Malay Penin. 5: 79, 1925. 
Fig. 178 
DIAGNOSIS OF HOLOTYPE: Stem slender, just 
below the leaves 4 mm in diameter; leaves 
67-68 cm long, 16 mm wide, subcoriaceous, 
green above, below paler and apparently some- 
what glaucous, with a central furrow over the 
midrib, 2 -pleated, and towards the tip the 2 
pleats spinulose above, at midsection with 16-18 
parallel secondary veins in each half, these 
prominent throughout, below the tertiary cross 
veins visible, remote and forming elongate 
meshes, the blade ligulate, in outer (4 tapering 
