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gradually to the subulate, deltoid apex about 
3 cm long and 1.5 mm wide, the base amplexi- 
caul, unarmed, partly dark purple, beginning at 
4 cm up the margins with prickles 0.7- 1.5 mm 
long, 3-5 mm apart, subulate, ascending at 45°, 
the base thickened, pale; the midrib below un- 
armed to beyond the middle; at midsection the 
margins with prickles 0.7-0.9 mm long, 2-4 
mm apart, subulate, ascending; near the tip the 
margins and midrib below with subulate ser- 
rations 0.4-0. 6 mm long, 0.7-1 mm apart; pis- 
tillate inflorescence erect, bearing a single syn- 
carp; peduncle 10 cm long, 4-5 mm in di- 
ameter, 3-sided, bracteate; syncarp 4.5 cm long, 
3.1 cm in diameter, oblong-ellipsoid, bearing 
about 120 drupes, these 15-17 mm long, 3-4.5 
mm wide and thick, pale green, fusiform but 
the upper end contracted and more slender, 5-6- 
angled, the body 11-12 mm long; pileus 8-9 
mm long, the base 2.5-3 mm high, narrowly 
semiorbicular, with 5-6 sharp angles, these 
mostly single; style 5-5-5' mm long, sharply 
curved proximally, flattened and ridged below, 
pale, bony; stigma 3-4 mm long, distal, linear, 
running to the apex, brown, papillose; endocarp 
centering in lower but extending to the 
base, the walls 0.1 mm thick, cartilaginous, pale; 
seed 8-9 mm long, extending to the base; apical 
mesocarp spherical, filled with white, medullary 
membranes; basal mesocarp sparse, fibrous. 
HOLOTYPE: Malaya, Kelantan, along the 
ridge of Gunong Sitong, 2,500 ft. alt., 6 March 
1924, Mo harried Nur & Foxworthy 12,252 
(sing). 
DISCUSSION: P. spinulosus is a member of 
the section Aero stigma, and in that section is 
found its closest relative, P. collinus Ridl., which 
species has the leaves 25-45 cm long, 8-11 mm 
wide, unarmed above, each half at midsection 
with 10-11 secondary parallel veins, at mid- 
section the margins with subulate-serrations 
0.2-0. 3 mm long; syncarp 3 cm long; drupes 
14-15 mm long, the pileus base hemispheric, 
abruptly contracted to the subulate style which 
is 4-5 mm long. P. spinulosus has the leaves 
67-68 cm long, 16 mm wide, above towards the 
tip the two pleats spinulose, at midsection with 
16-18 secondary parallel veins in each half, at 
midsection the margins with subulate prickles 
0.7-0.9 mm long; syncarp 4.5 cm long; drupes 
15-17 mm long, the pileus base narrowly hemi- 
spheric, gradually tapering into the thick subu- 
late style which is 5-5.5 mm long. 
Pandanus unguiculatus Ridl., Mat. FI. Malay 
Penin. 2: 229, 1907; FI. Malay Penin. 
5: 78, 1925; (sect. Aero stigma) 
Fig. 179 
DIAGNOSIS OF HOLOTYPE: "Plant small”; 
leaves 26-48.5 cm long, near the base 2-2.3 
cm wide, near the middle 2. 4-2. 7 cm wide, 
ligulate, but perceptibly tapering both ways 
from the middle, dark green above, pale green 
below, thick chartaceous, only the midrib thick- 
ened, but the blade pleated and M-shaped in 
section, the secondary parallel veins conspicu- 
ous, at midsection 17-19 in each side, the ter- 
tiary cross veins visible in upper third below, 
oblique, forming short, rather oblong meshes; 
near the apex abruptly contracted to a subulate 
tip more than 20-25 mm long, 0.9 mm wide, 
the base amplexicaul and entire but beginning 
at 4-5 cm the margins with prickles 0.7-1 mm 
long, 2-6 mm apart, subulate, ascending, pale; 
the midrib unarmed for lower 34 s; at the mid- 
section the margins with prickles 0.5-0.7 mm 
long, 2-5 mm apart, subulate, appressed as- 
cending, "red-tipped” but when dried brownish; 
near the apex the margins and midrib below 
with subulate-tipped serrulations 0.2-0. 3 mm 
long, 0.5-2 mm apart; peduncle 5 cm or more 
long, 4 mm in diameter, bracted; syncarp sin- 
gle, 5.5 cm long, 4 cm in diameter, broad ellips- 
oid, bearing about 220 drupes, these 16-18 mm 
long (or 18-20 mm, following the curve), 4-5 
mm wide, 3.5-4 mm thick, oblance-fusiform, 
the body 14-15 mm long, narrowly ellipsoid; 
the pileus smooth, pyramidal-ovoid, 5-7 mm 
high, tapering into the hornlike style which is 
2.5-3 mm long, mostly upwards curved to about 
right angles; stigma 2. 5-3. 5 mm long, distal, 
linear, papillose, brown; endocarp slightly sub- 
median, 9—1 1 mm long, oblanceoloid, the apex 
truncate, but with ascending shoulders partly 
surrounding the apical cavity, stramineous, carti- 
laginous, the lateral walls 0.15 mm thick; seed 
7-8 mm long; apical mesocarp forming a 
broadly ovoid cavern with a few white medul- 
lary membranes; basal mesocarp scant, fibrous 
and fleshy. 
