Page 290: Revision of Pandcmus, 15. Malaya — St. John 
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spike 14 cm long, bearing 4 syncarps; stigmas 
4 mm long, sharply proximally bent, the recep- 
tive surface usually proximal. 
HOLOTYPE: Malaya, Kelantan, Sungei Ketah, 
Batu Bau (Bow), 15 Feb. 1924, Mo h anted Nur 
& F oxworthy 12,104 (SING). Holotype exam- 
ined! 
SPECIMENS EXAMINED: Malaya, Pahang, Gua 
Tipus, the common pandan on the limestone, 17 
Oct. 1927, M. R. Henderson 19,468 (sing). 
DISCUSSION: P. irregularis was placed by Dr. 
Ridley in the section Rykia. It has, however, the 
drupes 1-3 -celled, and usually 2 -celled, with 
the cells in linear arrangement, and the large 
stigmas terminal, but distal and laterally di- 
rected. There is no question but that it must 
now be placed in the section Hombronia. 
It is one of the numerous plant species known 
to occur only on limestone knobs. These pre- 
cipitous pinnacles or narrow ridges rise spec- 
tacularly from the rolling uplands of Thailand 
and Malaya. Their sides are often precipices of 
shining, white limestone, sparsely vegetated, but 
their crests have a dark evergreen, virgin forest 
cover. To a botanist they are very attractive. 
Four species of Pandanus in Malaya are re- 
stricted to these calcareous habitats. Besides this 
species published by Ridley, two more are now 
being described, and one more has yet to be 
found in fruit. A good account of this localized 
flora is given by Dr. M. R. Henderson (Roy. 
Asiat. Soc., Malay Branch, Jour. 17: 13-87, pi. 
Ill — XI, 1939). 
Malaya, sect. nov. 
Phalangibus pluricarpellatis, stylis terminali- 
bus adscendentibus vel arcuatis corniformatis in- 
tegris vel bifidis, stigmatibus centripetalibus el- 
lipsoideis vel lanceolatis; plantis masculis in- 
cognitis. 
Phalanges of several united carpels; styles ter- 
minal, ascending or arching, stout, hornlike with 
a subulate apex or compressed and more or less 
oblong, with the apex sharply bifurcate; stig- 
mas centripetal, ellipsoid or lanceolate; stami- 
na te plants unknown. 
HOLOTYPUS: Pandanus Klossii Ridl., Fed. 
Malay States Mus., Jour. 6: 190-191, 1915. 
DISCUSSION : The single known species occurs 
on the mountains of Pahang in Malaya. It com- 
bines the characteristics of the section Rykia 
in its hornlike, simple or bifurcate styles, and 
of the section Pandanus, in its many-carpellate, 
large phalanges. Since these two sections are 
very distinct and are only remotely related, it 
is best to make the very bizarre P. Klossii the 
basis of a new section. 2 
Pandanus Klossii Ridl., Fed. Malay States Mus., 
Jour. 6: 190-191, 1915; FI. Malay Penin. 
5: 74, 1925; (sect. Malaya) 
Figs. 184, 185 
DIAGNOSIS OF HOLOTYPE: Shrub 2-7 m tall; 
stems usually solitary, 7.5 cm in diameter, gray, 
erect or arcuate; leaves 1.3-1. 8 m long, 6.8-72 
cm wide, thick coriaceous, above dark green, 
below pale green, 1 -ribbed, but 2-pleated and 
in section low M-shaped, the longitudinal paral- 
lel secondary veins 0.5-0.8 mm apart, 49-52 
in each half, evident on both sides as are the 
tertiary cross veins that form a network of 
meshes, oblong or rhombic, longer than broad, 
the blade ligulate, and only near the tip rather 
abruptly narrowing to a 5 cm stout subulate tip, 
trigonous and 2-3 mm wide, the very base not 
collected, near the base the margins with stout 
thorns 5-8 mm long, 8-30 mm apart, arcuate 
ascending, black; the midrib below with similar 
but reflexed thorns; at midsection the margins 
with prickles 3 mm long, 15-50 mm apart, 
subulate, ascending, flat appressed; the midrib 
below with prickles 2 mm long, 35-50 mm 
apart, heavy, arcuate, ascending; near the tip 
2 The new section can be inserted in the author’s key in this Revision of the Genus Pandanus (p. 2) by 
replacing the 11th line from the bottom by: 
L. Stigmas elongate, acerose, or stout hornlike, simple or bifurcate, 
a. Stigmas stout hornlike, simple or bifurcate Malaya 
a. Stigmas elongate, acerose, 
