Page 292: Revision of Pandanus, 15. Malaya — St. John 
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the margins and midrib below with serrations 
0. 3-0.5 mm long, those of the margin 1-1.5 mm 
apart, those of the midrib below 2-5 mm apart; 
pistillate peduncle 15-30 cm long, 12-18 mm 
in diameter, 3-sided, leafy bracted; syncarp sin- 
gle, globose or ellipsoid, about 15 cm in di- 
ameter; phalanges numerous and 6-6.5 cm long, 
3. 6- 3. 9 cm wide, 2-2.7 cm thick, cuneiform, 
4-6-angled, "showing signs of turning red or 
orange,’’ the sides smooth, gently convex, free 
in upper J4, the apex low convex, lateral sutures 
visible only in upper Vy central apical sinuses 
1- 4 mm deep, straight or arcuate, narrow; car- 
pels 9-10 (in a double 16-17 and the phalanges 
5.7- 6 cm wide), the apices semiorbicular , 
faintly angled; styles 4-6 mm long, dark, bony, 
either simple and hornlike or flattened and 
sharply bifid, mostly centripetal; stigmas as long 
as the styles, ellipsoid or lanceolate, brown, pap- 
illose, covering the proximal face; proximal 
sinus none; endocarp in lower 14 and 2.7 cm 
long, bony, massive, dark brown, the lateral 
walls 2 mm thick, the inner surfaces shining 
and rugulose; seeds 14-16 mm long, 2-3 mm 
in diameter, oblong-ellipsoid; upper mesocarp 
2- 2.5 cm long, of dense pith and a few fibers; 
basal mesocarp fibrous and fleshy. 
ADDED DESCRIPTION FROM ALL SPECIMENS 
EXAMINED: Phalanges about 96; carpels 5-10; 
syncarp 9-15 cm in diameter. 
HOLOTYPE: Malaya, Pahang, Gunong Tahan, 
July 1911, H. N. Ridley 16,037 (SING). Iso- 
type (bo). Ridley stated, "Common all over 
the Padang. In the more open exposed spots 
the stem is short and erect, about 6 to 8 feet 
tall; in the woods the stems are long and weaker, 
often falling about at all angles, 20 feet or 
more long. No trace of male flowers could be 
seen anywhere. The plant is very abundant, al- 
most filling up the woods in some places." The 
collection, Ridley 16,037, was made by Ridley 
previous to his publication of the species in 
1915, was labeled P. Klossii by him, and seems 
to have been the only collection in the Singa- 
pore herbarium up to that time. In his Flora 
of the Malay Peninsula (1925) he accepted the 
species and cited only his collection from Gu- 
nong Tahan, so this one seems acceptable as 
his holotype. 
SPECIMENS EXAMINED: Malaya, Pahang, 
Fig. 185. Pandanus Klossii Ridl., from holotype, 
syncarp, X Vl- 
Gunong Tahan, 14 Sept. 1937, E. J. H. Corner 
(SING) ; Pahang, Gunong Tapis, Kuantan, 4,600 
ft. alt., 14 June 1934, C. F. Symington & Kiah 
28,843 (sing). 
DISCUSSION: P. Klossii Ridl, a very unique 
species, is here made the holotype for the new 
section Malaya. This combines the characters of 
the two sections Pandanus and Rykia. When 
Ridley described the species in 1915 he did not 
place it in a section, but later in 1925 in his 
Flora of the Malay Peninsula he put it in the 
section Keura, which is now called Pandanus. 
As usual he had a good eye for species, but his 
description seems to have been hastily drawn. 
He stated ( 1915: 191 ) "the drupes are separate 
