350 
PACIFIC SCIENCE, Vol. XVII, July 1963 
till the fruit is nearly ripe, when from 6 to 7 
become adnate and remain so as the whole fruit 
breaks up.” The holotypic collection consists of 
two sheets, one with overripe fruit, the other 
with a small, immature syncarp. In both the 
phalanges are of several fused carpels. The car- 
pels are fused from the base to within 4 mm 
of the apex, and the sides lack sutures.The writer 
sees no evidence on these specimens to confirm 
Ridley’s statement that the drupes are separate 
and only tardily connate; on the contrary, all 
the structural evidence indicates that the pha- 
langes are of several carpels united from the 
beginning. 
Though Ridley did not explain the source 
of his specific epithet, it was obviously given in 
honor of Cecil Boden Kloss, an English zoolo- 
gist and to a lesser degree a botanist. In 1903 
and 1907 he was employed by the Singapore 
Botanic Garden. He was later director of the 
Raffles Museum. 
Multidens, sect. nov. 
Frutices, foliis angustis, syncarpio solitario el- 
lipsoideo, stylo singulo terminali osseoso prox- 
ime curvato basi cylindrico margine (2-) 3-6- 
lobato lobis deltoideis, drupis 1-loculatis. Plantis 
masculis cum inflorescentia bracteata 1-spicata, 
staminibus distinctis in axili affixis, filamentis 
4-plo tarn longis quam antheris. 
Bushes; leaves narrow; syncarp solitary, el- 
lipsoid; style single, terminal, proximally curved, 
bony, shining, the base cylindric, the margin 
(2-) 3-6-lobed, the lobes deltoid; drupes 1- 
celled. Staminate plants with inflorescence brac- 
teate, bearing a solitary terminal spike; stamens 
separate, attached directly to the axis; filament 
4 times as long as the anthers. 
HOLOTYPUS: P. pentodon Ridl., FI. Malay 
Penin. 5: 75, 1925. Also in the section is P. 
tetrodon Ridl. of Singapore. 
Pandanus pentodon Ridl., FI. Malay Penin. 5: 
75, 1925; (sect. Multidens) 
Fig. 186 
diagnosis OF lectotype: Stem short, 
branched, up to 2 cm in diameter; leaves said 
to attain more than 100 cm in length, 11-12 
mm wide, coriaceous, dark glossy green, pale 
green beneath, channeled above the midrib and 
with 2 lateral pleats, in section low M-shaped, 
narrowly ligulate, outer halves of leaves missing, 
at midsection with 17-22 secondary parallel 
veins in each half, the tertiary cross veins barely 
visible below near the base, the base amplexi- 
caul and unarmed, apparently copper-colored, 
beginning at 3-5 cm up the margins with pale 
serrulations 0. 5-0.8 mm long, 1-6 mm apart; 
the midrib unarmed to at least beyond the mid- 
dle; at midsection the margins with slender ser- 
rulations 0.3 -0.6 mm long, 4-8 mm apart, the 
upper surface unarmed; the apex tapering to a 
10 cm trigonous subulate tip which 10 cm 
down is 4 mm wide, the margins and midrib 
below with serrations 0. 2-0.3 mm long, 0.7-3 
mm apart; pistillate peduncle 13 cm long, 8 mm 
in diameter, bracted, bearing a single, erect 
syncarp; the highest peduncular bract 30 or 
more cm long, at base 2.2 cm wide, narrowly 
lanceolate and tapering noticeably right from 
the base, the margins with serrulations similar 
to those of the leaves, but on the upper surface 
beginning 10 cm up the 2 lateral pleats with 
similar serrulations, the lower ones 5-13 mm 
apart, the upper ones 2-5 mm apart; partly 
hiding the fruit are 3 series of floral bracts, 3 
deep, and subequal, 5 cm long, 2 cm wide, lance- 
elliptic, fleshy, cream-colored; syncarp 5 cm 
long, 3 cm in diameter, bearing about 400 
drupes (when young these are cream-colored), 
12-13 mm long, 2-3.5 mm wide and thick, 
linear-oblanceoloid, 5-6-angled, the body 10-11 
mm long, the summit rounded; pileus 2.5-3 
Insertion in Key 
On p. 225 (p. 2 of reprint) for the second J, read 
J. Styles 1 (-2), often cartilaginous or bony, 
z. Styles toothlike to broad subulate, entire or bifid; staminate spikes bearing stamens 
fascicled at apex of column Rykia 
z. Style with cylindric base, then curving proximally and the margin (2-) 3-6-lobed 
with deltoid lobes; staminate spike solitary, bearing separate stamens attached directly 
to the axis Multidens 
