Page 296: Revision of Pandanus, 15. Malaya — St. John 
353 
mm long, the base pyramidal-semiorbicular, 
smooth, when dried somewhat yellowish; style 
1-1.5 mm long, apical, bony, yellowish, shining, 
its base short cylindric, then the rest sharply 
curved proximally and flabellate with the mar- 
gins (3-) 4-5 (-6) dentate; stigma 0.5-1 mm 
long, obdeltoid to flabellate, papillose, brown, 
on the distal side of the style; endocarp central, 
bony, pale brown, the upper lateral walls 1 mm 
thick; seed 5-5.5 mm long; apical mesocarp 
an empty cavern; basal mesocarp fibrous up the 
sides, fleshy within. 
STAMINATE PLANT: Herbage not seen; stam- 
ina te inflorescence dense, leafy bracted, the main 
part 14 cm long; lowest floral bract about 30 
cm long, the upper part green, foliaceous, 10 
mm wide, with serrulations 0.2 mm long, 2-5 
mm apart, the 10 cm basal part 19-20 mm wide, 
folded, firm, apparently pale, with similar or 
smaller serrulations; median floral bract 7 cm 
long, 3 cm wide, navicular, elliptic, acute, ser- 
rulate near the apex on margins and midrib 
below, firm, apparently pale; spike single, ter- 
minal, 7.5 cm long, 13 mm in diameter, cylin- 
dric, dense; stamens distinct, attached directly 
to the axis; filaments 4.2-4.4 mm long, anthers 
0.8 mm long, narrowly oblong, bearing at tips 
a 0.15-0.2 mm subulate prolongation of the 
connective. 
LECTOTYPE: Malaya, Johore, Ulu Kahang, 
abundant along edges of Kahang river, 250 ft. 
alt., 1 June 1923, R. E. Holttum 10,861 (sing) 
— the pistillate plant here designated as lecto- 
type. Mingled on the same sheet is a staminate 
plant. 
Pandanus tetrodon Ridl., Roy. Asiat. Soc., Jour. 
68: 13-14, 1915; P. singaporensis Ka- 
nehira, Jour. Jap. Bot. 14: 173-177, figs. 
6-8, 10, 1938, a superfluous name; non 
Barrotia tetrodon Gaud., Bot. Voy La 
Bonite, t. 13, figs. 1-8, 1841, an invalid 
name; non P. tetrodon Balf. f., Linn. 
Soc. Bot. Jour. 17: 63, 1878, a name 
not accepted by its author; non P. tetro- 
don Balf. f. ex Kanehira, Bot. Mag. To- 
kyo 52: 236-239, fig. 70, 1938, a later 
synonym of P. compressus Mar tel li 
( 1905 ) ; ( sect. Multidens ) 
Fig. 187 
DIAGNOSIS OF HOLOTYPE: "Large bushy 
plant; branches at apex 2.5 cm in diameter”; 
leaves well over 97 cm long, 2.6 cm wide at 
the middle, 2 cm wide near the base, coriaceous, 
1 -ribbed, 2 -pleated, in section low M-shaped, 
about at the midsection with 25-26 secondary 
parallel veins in each half, these visible at least 
below throughout, tertiary cross veins visible in 
outer half, transverse, forming oblong meshes, 
mostly short, the blade ligulate, the apex not 
preserved, the base amplexicaul, unarmed, be- 
ginning at 5 cm up the margins with prickles 
1.5-2. 5 mm long, 5-10 mm apart, subulate- 
tipped serrae, pale, with reddish tips; the mid- 
rib below unarmed for at least lower %’s; near 
the midsection the margins with similar serrae, 
1.3-1. 5 mm long, 6-13 mm apart; leaf apex 
"acuminate,” but not preserved; pistillate in- 
florescence terminal, more or less concealed; pe- 
duncle 19 cm long, 7 mm in diameter, 3-sided, 
sinuous, leafy bracted, bearing 1 syncarp, this 
8.8 cm long, 5 cm in diameter, cylindric with 
obtuse ends, bearing about 176 drupes, these 
22-24 mm long, 6—10 mm wide, 5-10 mm 
thick, increasing upwards in size and the apical 
ones definitely the largest, cuneate-oblanceoloid, 
obtuse, 5-6-angled, upper % free, the sides 
nearly smooth; style 3-4 mm long, bony, dark 
shining brown, at base cylindric, then quadrate 
and curved proximally, and all the lateral ones 
with an oblique plane to the 2-3-4-dentate 
margins; stigma 1—2 mm wide or long, rugose, 
of various shapes, filling the cavity proximal 
of the style apex, with 2-4 (-5) deltoid lobes; 
endocarp centering in lower Vs, bony, dark 
brown; seed 6 mm long, obdeltoid; apical meso- 
carp 6-7 mm long, a rounded cavity with a few 
pale membranes; basal mesocarp fibrous and 
fleshy. 
EXPANDED DESCRIPTION OF PISTILLATE 
PLANTS: Stems at apex 1.5-2. 5 cm in diameter, 
brown, striate, covered with ascending adventi- 
tious rootlets 1-2 cm long; leaves 78 to more 
than 97 cm long, the tip gradually diminishing 
to a trigonous subulate apex about 10 cm long, 
and at base 4 mm wide, on the subulate apex 
the margins and midrib below with serrations 
0.3-0. 5 mm long, 2-5 mm apart; syncarp sin- 
gle or with a secondary one below and 4.5 cm 
long, ovoid. 
