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PACIFIC SCIENCE, Vol. XVII, October 1963 
lanceolate, tomentellous, often reddish within; 
stamens glabrous; ovary densely tawny or golden 
tomentellous; style hirsutulous; stigma dark 
maroon, glabrous; capsule commonly 16-34 mm 
broad, often subtended by the persistent sepals, 
the carpels with a dense persistent fulvous or 
reddish pilosity; endocarp firm, thickly pilose 
with pale hairs. 
TYPE: Hawaii: Kawaihae-iuka, in 1862, Wm. 
Hillebrand (in the Wawra Herbarium, Vienna). 
A photo shows capsules 27 mm broad. 
(a) var. hawaiensis 
The typical variety, which is certainly distinct 
from Pelea cinerea, as was stated (in herb.) by 
the late C. N. Forbes, is nonetheless a southeast 
counterpart of that species. The distribution of 
the typical variety includes Hawaii and Maui. 
Other varieties, some of them eminently distinct, 
others only poorly differentiated, are found on 
Hawaii, Maui, Lanai, and Molokai. 
( b ) var. rubra ( Rock ) B. C. Stone, comb. nov. 
P. cinerea var. rubra Rock in Bot. Gaz. 65: 
264, 1918. 
P. oblon^ifolia var. /3 Hillebrand, FI. Haw. 
Ids. 65, 1888. 
P. cinerea var. 8 Rock, Indig. Trees Haw. Ids. 
239, 1913. 
P. cinerea var. 8 Hillebrand, op. cit. 69. (as 
to Kau specimen). 
Hillebrand tells us that this is a glabrous 
shrub with long rambling branches. The poorly 
KEY TO VARIETIES OF Pelea hawaiensis 
Capsules mostly 27-34 mm diameter (rarely smaller); inflorescences often few-flowered (3-9). 
Leaves puberulent beneath, at least on the costa, at first ( and often persistently ) uniformly 
tomentellous to pilose; inflorescence compact, with stout axes, peduncles ca. 2 mm thick, 
densely tomentellous; capsular pubescence fulvous to brownish-orange. Hawaii 
var. hawaiensis 
Leaves glabrous, glossy; inflorescences elongate, the axes slender, peduncle to 1 mm thick, 
glabrous or nearly so; capsular pubescence reddish. Hawaii var. rubra 
Capsules smaller, mostly 16-22 mm diameter; inflorescences mostly compact and multiflorous 
with 9-27 flowers, or rarely more open and with fewer flowers. 
Blades densely shaggy-pilose beneath, short-lanceolate; branchlets puberulent; inflorescences 
mostly 3-9-flowered. 
Blades cordate at base; tomentum fulvous, the hairs extremely minute. Molokai 
var. molokaiana 
Blades subcuneate to rounded at base; tomentum brownish or olivaceous, the hairs up to 
0.7 mm long. Maui, Lanai var. pilosa 
Blades tomentellous to glabrate beneath, sometimes at last glabrous; inflorescence several- 
to many-flowered (with up to 27 flowers), or less commonly only 3-7 -flowered. 
Inflorescences mostly only 3-7-flowered. 
Blades cordate or emarginate at base, slightly puberulent on the costa dorsally, elon- 
gate and often acuminate. Lanai... var. sulfurea 
Blades cuneate at base, glabrate to glabrous beneath, rather broad, ovate. Maui 
var. Brighamii 
Inflorescences mostly 9-27-flowered. 
Blades cordate or subcordate at base, or sometimes merely emarginate; inflorescences 
about 4-6 cm long. 
Blades glabrate beneath; petioles mostly 20-45 mm long. Maui var. Remyana 
Blades puberulent beneath; petioles mostly 7-18 mm long. Maui ..var. racemiflora 
Blades cuneate or rarely slightly emarginate at base; inflorescences compact, 2-3 cm 
long; costa puberulent or glabrous dorsally. Hawaii var. Gaudichaudii 
