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PACIFIC SCIENCE, Vol. XVII, October 1963 
glabris, deltoideis, 7-9-nervatis; petalis toto gla- 
bris sparse obscure punctatis ovato-lanceolatis 
ca. 11 mm longis et 5 mm latis apice intus un- 
guiculatis; staminibus fertilibus 8, (4 ad 10 mm 
longis, 4 ad 7.5 mm longis, antheris 1.7— 1.8 
mm longis, filamentis ligulatis ca. 1 mm latis); 
ovario glabro ca. 2 mm alto, disco glabro ca. 
1 mm alto; stylo ca. 2 mm longo, stigma ca. 1.4 
mm lato lobis rufidis minute papillatis cruciatis; 
floris foemeneis minoris sepalis ca. 3-5 mm 
longis et 3 mm latis, petalis ca. 7-8 mm longis 
et 3.5 mm latis, staminibus sterilibus ca. 2 mm 
longis subaequitantibus; ovario discoque glabro, 
sytlo glabro ca. 4 mm longo; capsulis quadrilo- 
batis, glabris, ad 20-25 mm diametro, endo- 
carpio glabro. 
HOLOTYPE: Kauai: Kokee, August 1924, 
Degener 8586 (Bishop; duplicates at Cath, NY). 
DISTRIBUTION: Kauai, in the Waimea high- 
lands. 
SPECIMENS EXAMINED: Kauai: Near Lehua- 
makanoe Bog, 18 September 1955, Degeners, 
Hansen & Hanner 23984 (Bishop; staminate 
fls. ). Kaholuamano, October 1895, Heller 2869 
(formerly determined as P. resiniflora ) (GH, 
edinb, US). Halemano, 14 February 1909, Rock 
2294 (A). Waialeale, October 1911, Rock (a). 
Kaholuamano, September 1909, Rock 5551 (A, 
Bishop), 5552 (gh, Bishop). 
This is by no means a complete citation of 
specimens; more are extant, particularly in the 
Bishop Museum, but many lack data, particularly 
localities and collectors’ numbers. Most of these 
specimens had been determined as Pelea macro- 
pus Hillebr., which however is quite distinct in 
having puberulent ovaries ( and hence capsules ) 
and clavoid pedicels. Because of this interpreta- 
tion, this species has consistently been mistaken 
for P. macropus, and this error has tended to 
obscure knowledge of both of them. The gla- 
brous endocarp and rotate capsular lobes place 
P. Lohiauana in a group with P. oblanceolata 
St. John (of Hawaii), P. manukaensis St. John 
(of Hawaii), and P. pseudoanisata Rock (of 
Hawaii ) . 
The specific name honors Prince Lohiau of 
Kauai, hero of the tales of Pele and Hiiaka, and 
one of the memorable characters of Hawaiian 
mythology. 
8. Pelea macropus Hillebrand, FI. Haw. Ids. 65, 
1888. 
P. acutivalvata Leveille in Fedde Repert. Sp. 
Nov. 10:153, 1911. 
Innovations minutely and closely pale cinere- 
ous-puberulent, hairs appressed, ca. 0.2 mm 
long; branchlets at youngest nodes cinereous but 
soon glabrate; leaves glabrous ventrally, at first 
minutely cinereous dorsally (mostly on the 
costa ) , but soon glabrate, the older leaves mostly 
quite glabrous; petioles soon glabrate. Leaves 
opposite, the petioles mostly 9-15 mm long; 
blades thinly coriaceous, rounded or emarginate 
at apex, contracted and subcuneate at base (the 
margins sometimes slightly revolute); costa 
ventrally pale, shallow-sulcate; dorsally pale and 
prominent with a rather sparse and minute 
ephemeral puberulence; lateral nerves at right 
angles to costa or slightly ascending, about 7-9 
per side, conspicuous, prominulent on both sur- 
faces, connected at a distance of 2-10 mm from 
the margin by a moderately to deeply arcuate 
marginal nerve, with several sets of intervening 
meshes. Inflorescences small, axillary or ramu- 
line, mostly 1-3 -flowered, the peduncle very 
short, about 1-5 mm long, bibracteolate at apex, 
puberulent, the bractlets deltoid, rounded, 0.5 
mm long, puberulent; axes minutely cinereous, 
short ( to 5 mm ) in flower, somewhat elongated 
in fruit; pedicels 2-5 mm long, flared at base 
of calyx and clavoid, bibracteolate, cinereous; 
pistillate flowers with broadly deltoid sepals 
about 1.5 mm long and 1.7 mm broad, minutely 
puberulent externally; petals about 2.5 mm long 
and 1.7 mm broad, with a faint close puberu- 
lence of few minute hairs along the midregion, 
glabrous within; rudimentary stamens glabrous, 
less than 1 mm long; disk reddish, glabrous, 
shallow, ca. 1.7 mm broad; ovary puberulent, 
ca. 1 mm high and 1 mm broad; style glabrous, 
ca. 1.6 — 1.7 mm long (including stigma), stigma 
lobes red, each 0.25 mm long, minutely papillate, 
obovoid. Capsules 25-35 mm diameter, about 
9 mm high, sparsely appressed-puberulent (es- 
pecially near the base), the lobes deeply separated 
( Y <$ ) , somewhat acuminate and laterally com- 
pressed, often unequally developed; endocarp 
glabrous, cartilaginous; seeds 5-6 mm long, 
ovoid, the testa crustaceous, blackish. 
