Critical Species of Pelea — Stone 
417 
HOLOTYPE: Oahu: Koolau Range; Niu, Oc- 
tober 1940, St. John 20111 (Bishop). 
DISTRIBUTION: Endemic to the southeastern- 
most part of the Koolau Range. 
This variety differs from the typical P. pedun- 
cularis only in the much narrower leaves and 
perhaps slightly fewer-flowered cymes. 
SPECIMEN EXAMINED: Oahu: Northeast slope 
of Puu Kumakalii, April 1936, Degener, Tam, 
Takamoto & Martinez 10579 (Bishop, Catholic, 
NY). 
(c) var. pauciflora (St. John) B. C. Stone, 
comb. nov. 
Pelea Rockii var. pauciflora St. John in 
Lloydia 7:271, 1944. 
Cymes mostly 3-7-flowered. 
HOLOTYPE: Oahu: Koolau Range; Kaukona- 
hua Gulch, Wahiawa, May 1909, Rock 3046 
(Bishop). 
DISTRIBUTION: Known only from the type 
locality. 
A poorly defined variety which may be only 
an anomalous form of the species. 
(d) var. paloloensis (St. John) B. C. Stone, 
comb. nov. 
Pelea paloloensis St. John in Lloydia 7:271, 
1944. 
Leaves whorled in fours at the nodes. 
HOLOTYPE: Oahu: Koolau Range; Palolo, 
Waialae-iki, Jan. 1911 , Forbes 2404.0. (Bishop). 
DISTRIBUTION: Known only from the type 
locality. 
The variety, which probably is nothing more 
than an anomalous form, is an example of the 
occasional exception to the common taxonomic 
criterion for distinguishing species which are 
members of the section Pelea, with ordinarily 
whorled leaves, and the other sections, with 
generally opposite leaves. 
(e) var. quadrata B. C. Stone, var. nov. 
Fig. 4 
Capsulis elobatis quadratis fere ca. 23 mm 
latis. 
HOLOTYPE: Oahu: Waianae Mountains; Puu 
Hapapa, August 1932, Degener 8521 (Bishop). 
DISTRIBUTION: Known from the northern 
end of both ranges of Oahu. 
A characteristic variety, found both in the 
northern Waianae mountains and in the north- 
ern Koolau Range (as for instance around Pu- 
pukea), sometimes occurring in company with 
the typical variety. 
10. Pelea olowaluensis St. John in Lloydia 7:2 66. 
1944. 
A tree up to 3 m high, the branchlets at first 
puberulent; leaves opposite; petioles glabrous; 
blades 2.8-8 cm long, oval, subcoriaceous, gla- 
brous; cymes mostly 3 -flowered, glabrous, the 
peduncle ca. 9 mm long; capsules 16-20 mm 
broad, flattened, lobed halfway or less, glabrous; 
endocarp glabrous. 
HOLOTYPE: Maui: Olowalu Valley, on the cen- 
tral ridge, May 1920, Forbes 2326.M. (Bishop). 
DISTRIBUTION: West and East Maui. 
SPECIMEN EXAMINED: West Maui: Mauna 
Huuma, May 1910, Forbes & Cooke 22.M. 
(Bishop) . 
This species is closely allied to Pelea pedun - 
cularis of Oahu and falls properly into section 
Cubicarpa . The characteristically revolute basal 
leaf-margins are especially noticeable. When 
fresh the capsules might perhaps be unlobed or 
barely notched. The disjunction of the two col- 
lections would imply that the species is, or was, 
fairly widespread on Maui. 
Fig. 4. Pelea peduncularis var. quadrata Stone. 
Three views of a capsule in the fresh state, from 
Stone 2832 ( Pupukea, Oahu ) . 
