Revision of the Genus Pandanus Stickman, Part 16 
Species Discovered in Thailand and Vietnam 
Harold St. John 1 
The following new species are all but one 
based upon specimens collected by others and 
preserved in Bangkok, either in the herbarium 
of the Department of Agriculture at Kasetsart 
University or in the herbarium of the Royal 
Forestry Department. 
SECTION Aero stigma 
Pandanus biplicatus sp. nov. (sect. Aero stigma) 
Fig. 192 
DIAGNOSIS HOLOTYPI: Licet acaulescens, foliis 
1.01 m longis 3.6 cm latis parte l A basali sub- 
coriacea parte ultima crassiter chartacea supra 
viridibus infra pallidis et licet glaucis 1-sulcatis 
acute 2-plicatis in sectione mediali M-formatis 
et cum 41 nervis parallelis secundariis in quoque 
dimidio, nervis tertilis in parte 2 A ultima con- 
spicuis et reticulis anguste oblongis formantibus, 
laminis ligulatis subiter in apice trigono subu- 
late 3 cm longo 1-2 mm lato diminuentibus 
basi amplexicauli et inermi sed ex 14-15 cm 
marginibus cum aculeis 1.5-2 mm longis 2-8 
mm separatis subulato-serratis complanatis lu- 
teis, midnervo infra ex 20 cm cum aculeis 2-3.5 
mm longis 8-15 mm separatis subulatis reflexis, 
in sectione mediali marginibus cum aculeis 
1.5-2 mm longis 5-10 mm separatis subulatis 
proxime adscendentibus, midnervo infra cum 
aculeis 2.5-3 mm longis 15—22 mm separatis 
subulatis adscendentibus, in apice subulato mar- 
ginibus et midnervo infra cum serrulis 0.2-0.3 
mm longis 1.5-3 mm separatis, in pagina superi- 
ori ad apicem plicis cum serrulis 0.2-0. 3 mm 
longis 1-4 mm separatis, inflorescentia foeminea 
erecta et recta, pedunculo 22-30 cm longo gracili 
clavato 9-12 mm diametro in apice folioso brac- 
teato, bracteis herbaceis coloratis sed in sicco 
1 B. P. Bishop Museum, Honolulu 17, Hawaii, 
U.S.A. Manuscript received September 28, 1961. 
nigro-brunneis infima 19 cm longa 23 mm lata 
lineari-lanceolata ad basem integra parte X A ul- 
tima proxime subulato-serrata, syncarpio 9.5 cm 
longo 7.2 cm diametro obovoideo-ellipsoideo 
cum drupis numerosis, eis 32-34 mm longis 
6-7.5 mm latis 4-6 mm crassis anguste oblanceo- 
fusiformibus 1-carpelatis parte l A supera libera 
corpore 24-26 mm longo oblanceoloideo lateri- 
bus planis, pileo 13-14 mm longo cartilagineo 
basi 5-6 mm alta rotundato-pyramidali 6-angu- 
lata, stylo 6-8 mm longo crassiter subulato 
anguloso divergent! subsinuoso, stigmate 3-4 
mm longo distali lineari brunneo sulcato papil- 
loso in apice extenso, endocarpio in parte }A> 
infera obovoideo cartilagineo stramineo in apice 
truncato lateribus 0.1 mm crassis, semine 8-9 
mm longo obovoideo, mesocarpio apicali 7 mm 
longo cum medulla alba delicata completa, meso- 
carpio basali 5-6 mm longo fibroso et carnoso. 
diagnosis of holotype: Perhaps acaules- 
cent; leaves about 1.01 m long, 3.6 cm wide, 
the lower l A subcoriaceous, the outer 2 A thick 
chartaceous, green above, pale and apparently 
glaucous below, 1 -furrowed above the midrib, 
sharply 2 -pleated, at midsection M-shaped in 
section and with 41 parallel secondary veins in 
each half, tertiary cross veins conspicuous in 
outer 2 A’s , forming narrow oblong meshes, the 
blade ligulate, abruptly narrowed to a trigonous 
subulate apex 3 cm long, 1-2 mm wide, the 
base amplexicaul and unarmed, but beginning at 
14-15 cm the margins with prickles 1.5-2 mm 
long, 2-8 mm apart, subulate-serrate, flat, yel- 
lowish; the midrib below beginning at 20 cm 
with prickles 2-3.5 mm long, 8-15 mm apart, 
subulate, reflexed; at midsection the margins 
with prickles 1.5-2 mm long, 5-10 mm apart, 
subulate, closely ascending; the midrib below 
with prickles 2.5-3 mm long, 15-22 mm apart, 
subulate, ascending; on the subulate apex the 
margins and midrib below with serrulations 
0.2-0. 3 mm long, 1.5-3 mm apart; on upper 
466 
