Page 309: Revision of Pandanus, 16. Thailand, Vietnam— St. John 
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rowed over the midrib, 2 -pleated, chartaceous, 
green above, paler green below, ligulate, taper- 
ing to a 9 cm subulate, trigonous apex 0.3-0.8 
mm wide, at midsection with 2,6 -29 secondary 
parallel veins in each half, the base unarmed, 
but beginning at 4. 5-6, 5 cm the margins with 
prickles 1.3-2 mm long, 3-7 mm apart, subulate, 
ascending, yellowish; the midrib below begin- 
ning 6-8.5 cm up with prickles 1.5-2. 5 mm 
long, 6-15 mm apart, stout subulate, fully re- 
flexed arcuate; at midsection the margins with 
serrulations 0.2-0.6 mm long, 3-7 mm apart, 
the midrib below with remote prickles 2 mm 
long, subulate, arcuate, reflexed or ascending; on 
the subulate tip the margins and midrib below 
with serrulations 0.2 mm long, 0.5-2 mm apart; 
pistillate inflorescence terminal, erect, with 1 
syncarp or rarely with a small secondary one; 
peduncle 15 cm long, clavate, trigonous, 5 mm 
in diameter at apex, with a few leafy bracts on 
upper half, the upper bracts 40-50 cm long, 
15-18 mm wide; syncarp 2. 5-4.5 cm in diame- 
ter, globose to ellipsoid, green, bearing about 
160 drapes, these 15-18 mm long, 4-5 mm 
wide, 3-4 mm thick, subfusiform, upper 2 A free, 
the body oblance-ellipsoid, obtuse, 10-12 mm 
long; pileus 9-11 mm long, the base 2-3 mm 
high, oblate pyramidal-hemispheric, smooth, 
5-7-angled, the angles continuing well up the 
style; style 5-7 mm long, stout, 5-7-angled, su- 
bulate, proximally arcuate, the outer part bony, 
shining, yellowish; stigma 2.5-3 mm long, distal, 
sublinear, creased, brown, papillose, running al- 
most to the apex; endocarp in lower 14, carti- 
laginous, yellowish, the walls 0.1 mm thick, the 
inside shining; seed 6-7 mm long, 2-2.5 mm in 
diameter, oblanceoloid, truncate; apical meso- 
carp 2 mm long, hemispheric or depressedly so, 
containing white, medullary membranes; basal 
mesocarp fibrous and fleshy. 
STAMINATE PLANTS: Prostrate, stems 7-11 
mm in diameter, very short; leaves 80-90 cm 
long, 18 mm wide at the middle and base, the 
subulate apex 3-4 cm long, at midsection with 
24-27 parallel secondary veins in each half; 
staminate inflorescence almost erect, 18 cm long, 
fragrant; peduncle 7 cm long; flowering part 
11 cm long, with conspicuous yellow bracts, the 
lowest one 9.5 cm long, 16 mm wide, linear 
lanceolate, chartaceous, serrulate only on the 
short subulate tip; median bract 6 cm long, 14 
mm wide, lanceolate, the margins of the outer 
half and the midrib below near the tip with 
pale, fragile serrulations 0.1 mm long; spikes 
about 7 and 1.5-2 cm long, 8-15 mm in diame- 
ter, dense; stamens numerous, attached singly, 
directly to the axis; filaments 0. 1-0.2 mm long; 
anthers 6-7 mm long, linear, bearing at the apex 
a projection of the connective 0.2-0.3 mm long, 
sometimes subulate but more commonly flat, 
lanceolate, obtuse. 
HOLOTYPUS: Thailand, Peninsular, Krabi, 83 
km. from Huay Yawt, very common in moist 
localities by roadside, 100 m. alt., 28 Jan. 1958, 
T em Smitinand 4,1 43 ( BKF ) . 
SPECIMENS EXAMINED: Thailand, Trat, Koh 
Chang, shrub in evergreen forest by stream, 40 
m. alt., 19 Feb. 1954, ( BKF ) ; erect shrub 40 cm. 
tall, common in swampy ground in evergreen 
forest, 1 June 1952, Bunnak ( Sang Kachand ) 
333 (BKF); Trat, Huayraent, Tern Smitinand 
1,392 (BKF); Peninsula, Trang, Kantang, Khuan 
nang hong, less than 100 m. alt., common in 
evergreen jungle, 26 Aug. 1955, Tem Smitinand 
3,012 (BKF); S. E. Part, Trat, Kaw Kut, Ao 
Salat, common in evergreen forests, 5 m. alt., 5 
April 1959, Tem Smitinand 3,702 , staminate, 
( BKF ) ; S. E. Part, Trat, Huang Raeng, Dong 
Tapit, prostrate, common in swamp by path in 
evergreen jungle, under 50 m. alt., 21 June 1952, 
Tem Smitinand 1,391, staminate, (bkf no. 
7,276); Kasvon, Nov. 1896, C. Curtis 3,247 
(SING). 
MALAYA: Penang, near Tanjong Toking, July 
1889, C. Curtis 1,821 (SING); Perak, Lahat, 
Ipoh, Oct. 1904, H. K.^Riidley'], (sing),, one 
of the original paratypes of P. perakensis RidL 
DISCUSSION: P. Toei is a member of the sec- 
tion Aero stigma, as is its closest relative, P, Kin- 
gianus Mar tell i of Malaya, a species which has 
the drupes 11-13 mm long; style 5 mm long; 
pileus rounded; and the leaves 40-50 cm long, 
14 mm wide. P. Toei has the drupes 15-18 mm 
long; style 5-7 mm long; pileus prominently 
angled; and the leaves 96-130 cm long, 20 mm 
wide. 
The new epithet is the vernacular name of 
the plant in its native region in Thailand. 
