Page 313: Revision of Pandanus , 16. Thailand, Vietnam- St. John 
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endocarp inframedian, lacking distinct marginal 
flanges; bark grayish, unarmed; and the leaves 
5 m long, 8-20 cm wide, the apex abruptly short 
acuminate. P. calcis has the syncarp 6-7 cm in 
diameter, subglobose, drupes 3. 2=3. 8 cm long, 
not viscid, the upper Vl free; endocarp supra- 
median, with large lateral flanges; bark armed 
with short spines; and the leaves 75-84 cm 
long, 2=2.7 cm wide, gradually long tapering to 
the subulate apex. 
The new epithet is from the Latin, calx , lime, 
and is given in reference to the habitat of the 
species, the side cliffs or summits of precipitous 
limestone ridges. 
SECTION Rykia 
Pandanus bifidus sp. nov. (sect. Rykia) 
Fig. 196 
nom. vern.: "t6i noi.” 
DIAGNOSIS HOLOTYPI: Planta subacaulis, fo- 
lds 1.3 m longis 3.3 cm latis subcoriaceis 
gladiatis in sectione mediali cum 32 nervis 
secundariis parallelis in quoque latere in apice 
subulato breve diminuentibus et 10 cm ex apice 
14 mm latis proxima basem marginibus cum 
aculeis 3.5=4 mm longis 25-35 mm distantibus 
subulatis adpressi-adscendentibus stramineis, 
midnervo infra cum aculeis 2.5-3 mm longis 
50-58 mm distantibus simulantibus sed valde 
reflexis, in sectio mediali ; marginibus et mid- 
nervo infra cum aculeis 2.5-3 mm longis 15-48 
mm distantibus adscendentibus, proxima apicern 
marginibus et midnervo infra cum serris 0.8-1. 5 
mm longis 2=7 mm distantibus, syncarpio soli- 
tario terminal! 12 cm longo 10 cm diametro ap- 
parente ellipsoideo cum drupis numerosis eis 
3-3.2 cm longis 8-10 mm latis 8-11 mm crassis 
oblongo-oblanceoloideis 5-7-angulosis lateribus 
subplanatis, parte Va supera libera, pileo 4-5 
mm alto semiorbiculari 5-7-anguloso sublaevi 
tarde deciduo, stylo 3.5-5 mm longo valido 
osseoso corniformato eis lateralibus proximo- 
curvatis irregulariter bifidis vel in eis apicalibus 
integris et subulatis, stigma d 3-4 mm longo 
brunneo papilloso proximo lanceolate vel bifido, 
endocarpio mediali osseoso obovoideo acumi- 
nate pallido sed intra obscure brunneo lucido 
lateribus 1 mm crassis, seminibus 12 mm longis 
obovoideis, mesocarpio apicale cavernoso, meso- 
carpio basali fibroso et carnoso. 
DESCRIPTION OF ALL SPECIMENS EXAMINED: 
Trunk from almost none to 3 m tall; leaves 
1.3-3 m long, 3.3-5 cm wide, subcoriaceous, 
gladiate, at midsection with 32 secondary veins 
in each half, tapering to a short subulate apex 
which 10 cm down is 14 mm wide, near the 
base the margins with prickles 3.5-4 mm long, 
20-35 mm apart, subulate, appressed ascending, 
stramineous; the nearby midrib below with 
prickles 2.5-3 mm long, 50-58 mm apart, 
similar but sharply reflexed; at the middle the 
margins and midrib below with prickles 2.5-3 
mm long, 15-48 mm apart, ascending; near 
the apex below finely reticulate, and the margins 
and midrib below with serrae 0.8-2 mm long, 
2-17 mm apart; syncarp solitary, terminal, 12 
cm long, 10 cm in diameter, apparently ellip- 
soid, bearing very numerous drupes, these 3-3.2 
cm long, 8-10 mm wide, 8-11 mm thick, ob- 
long-oblanceoloid, 5-7-angled, the sides almost 
plane, upper Va free; pileus 4-5 mm high, semi- 
orbicular, 5-7-angled, nearly smooth, finally de- 
ciduous; style 3.5-5 mm long, stout, bony, horn- 
like, the lateral ones sharply curved towards 
apex of syncarp, irregularly bifid, or on the 
apical ones entire and subulate; stigma 3-4 mm 
long, brown, papillose, proximal, lanceolate or 
bifid; endocarp median, bony, obovoid, with an 
apical acumination, pale except for the dark 
brown, shiny inner surface, the lateral walls 1 
mm thick; seed 12 mm long, obovoid; apical 
mesocarp cavernous; basal mesocarp fibrous and 
fleshy. 
HOLOTYPUS: Thailand (Siam): Pak Tong 
Chai, Nakawn Rachsima (=Korat), marshy 
ground, 200 m. alt., Dec. 27, 1923, A. F. G. Kerr 
8,132 (bk). 
SPECIMENS EXAMINED: Thailand (Siam), 
Kao Krading, Loi, evergreen forest, 1,200 m. 
alt., Feb. 11, 1931, A. F. G. Kerr 20,084 (bk). 
DISCUSSION : P. bifidus St. John is a member 
of the section Rykia. Its closest relative is P. 
furcatellus Martelli from Tonkin. This has the 
endocarp in the lower Vdl apical mesocarp cav- 
ernous, 10-11 mm long; and the style forks 
ovate, acute. In contrast P. bifdus has the endo- 
carp median; apical mesocarp cavernous, 4 mm 
long; and the style forks subulate. 
The new epithet is the Latin, bifdus, two- 
cleft, in reference to the cleft style. 
