50 
Fig. 5. Male claspers, elegantulus group. Felhacus 
elegantuhis (Reuter): a. Right clasper (posteroventral); 
b, left clasper (lateral). Felisacus ochraceus Usinger: 
c, Right clasper (posteroventral) ; d, left clasper (lateral). 
If critical study of a more complete series 
of species should uphold this or a similar 
division of the genus, it might be found ad- 
vantageous to regard the groups as subgenera. 
In this case, provided elegantulus did not 
prove to fall into the same group as glabratus, 
Hyaloscytus Reuter, 1905, could be retained as 
a subgeneric name for the elegantulus group. 
Each of the groups as tentatively desig- 
nated above centres on a more or less distinct 
geographical range, with a number of aber- 
rancies to be expected as the result of dis- 
persal from the centre of origin. The first 
group is mainly northern Pacific, the second 
mainly southern Pacific, and the third more 
or less fringing the Indian Ocean. 
It seems to the author not unlikely that 
some of the described species of Felisacus 
might eventually have to be considered as 
subspecies. For example, fillclcola is very close 
to elegantulus and might have developed as 
a geographical race of the latter, whereas 
usingeri is close to philippinensis, and the for- 
mer might perhaps prove to be a lowland 
race and the latter a high-altitude race of the 
one bitypic species. Elowever, at present there 
are no criteria by which specific and sub- 
PACIFIC SCIENCE, Vol. VIII, January, 1954 
specific limits in this genus might be satis- 
factorily demonstrated; until more complete 
evidence is available on extent of variation, 
distribution, biology, and interbreeding, and 
the structure of the male genitalia, it seems 
desirable to maintain specific status for all 
forms which are readily distinguishable on 
the basis of correlated structural differences. 
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