Studies in Fruit Flies. Part I. Dacini — Hardy AND Adachi 
149 
Callantra Walker 
Callantra Walker, I860, Linn. Soc. London, 
Proc. 4: 154. 
Melksis Bezzi, 1916, BuL Ent, Res. 7: 114. 
Hendel, 1914, Wien Ent. Zeit. 33: 74. 
On the basis of the material which has 
been studied to date this appears to be a 
distinct genus. It can be distinguished from 
Dacus s. 1. by the following characters. The 
antennae are much more elongate, the second 
and third segments combined are about equal 
to the vertical length of the head (Fig. 2b) ; in 
Dacus the two segments combined are about 
0.5 or 0.6 as long as the head. In Callantra the 
first antennal segment is elongate and equal 
to the second and nearly equals the visible 
portion of the palpi ; in Dacus the first segment is 
much shorter than the second and is scarcely 
over 0.25 as long as the visible palpi. In 
Callantra the abdomen is strongly clavate and 
petiolate, and a prominent hump is present 
on each side of the petiole (base of first seg- 
ment). The flies are markedly wasp-like in 
appearance. The abdomen is strongly arched 
from a lateral view and is hollowed out be- 
neath. The ovipositor is cylindrical instead of 
flattened dorsoventrally, as is usual in Dacus. 
Callantra also has no prescutellar bristles and 
no supernumerary lobe in the wing of the 
male. ■ 
Malloch (1939^.' 410) considers this a sub- 
genus of Dacus, and it is entirely possible 
that, when the group has been studied in 
detail throughout its geographic range, most 
of the above characters will intergrade with 
Dacus. 
genotype: Callantra smieroides Walker. 
Nine species of Callantra are known from 
the regions covered in this study. Just two, 
however (C. smieroides Walker and C. longi- 
cornu (Wiedemann)), were represented in the 
collections studied, and as the original de- 
scriptions of some of the species do not give 
enough information it has not been possible 
to devise a satisfactory key to the species at 
this time. 
The following have also been recorded 
from these regions: 
C. axana Hering, 1938, Deut. Ent. Ztschr., 
410, pi. II, fig. 4. Type locality: Key Island. 
C bioculata (Bezzi), 1919, Philippine Jour. 
Sci. 15(5): 437-438, pl.II, fig. 4. Type locality: 
Luzon, Philippine Islands. 
C. conopsoides (de Meijere), 1911, Tijdschr. 
V. Ent. 54: 378-380. Type locality: Java. 
Misspelled '^Conopoides' by Bezzi, 1919, Phil- 
ippine Jour. Sci. 15(5): 438, 440. 
C nummularia (Bezzi), 1919, Philippine 
Jour. Sci. 15(5): 441-442, pi. II, fig. 6. Type 
locality: Luzon, Philippine Islands. 
C. pedunculata (Bezzi), 1919, Philippine 
Jour. Sci. 15(5): 439-440, pL II, fig. 5. Type 
locality: Luzon, Philippine Islands. 
C splendida Perkins, 1938, Roy. Soc. 
Queensland, Proc. 49: 136-137. Type locality: 
Java. 
C subsessilis (Bezzi), 1919, Roy. Soc. 
Queensland, Proc. 49: 435-436, pi. II, fig. 
3. Type locality: Panay, Philippine Islands. 
Callantra longicornis (Wiedemann) 
Fig. 1 
Dacus longicornis Wiedemann, 1830, Auss 
Zweifl. Ins. 2: 524. 
Bactrocera vespoides Doleschall, 1859, Na- 
tuurk. Tijdschr, Nederland. Indie 17: 123. 
This species fits in a distinct group of 
Callantra distinguished by the presence of a 
pair of strong spines on the underside of the 
front femora near their apices (Fig. id). We 
know of only one other named species (C. 
conopsoides (de Meijere)) which possesses this 
character. We do, however, have on hand an 
apparently unnamed species from India which 
has these spines. C. longicornis is closely re- 
lated to C. conopsoides (de Meijere), but it 
differs from de Meijere’s description as fol- 
lows: The legs are almost entirely yellow to 
rufous, not with the femora and tibiae chiefly 
blackish brown; the face has a moderately 
small brown to black spot in each antennal 
furrow, not with black markings on the oral 
