Studies in Fruit Flies. Part L Dacini — Hardy AND AdaCHI 
167 
tion of cell R3. The anterior sclerite of each 
propleuron, the coxae, and tibiae are usually 
discolored with brown to black as in pedestris, 
and the occiput is predominantly rufous as 
in typical dorsalis. The genitalia of both sexes 
appear to be identical with those of typical 
dorsalis. 
This is apparently the common form of the 
dorsalis complex which occurs in the Philip- 
pine Islands, and it was thought that it rep- 
resented a geographically distinct subspecies. 
It is now obvious that the ranges of typical 
dorsalis and dorsalis occipitalis overlap in Ma- 
laya, and on the basis of this geographic 
intergrading it is probably best that occipitalis 
be considered a variety. The extension of the 
costal band beneath vein R3 is somewhat 
variable and is much more distinct in some 
specimens than in others. It appears to be 
much more constant in specimens from the 
Philippines than in those from other areas. 
Length: 7. 0-8.0 mm. 
TYPE LOCALITY: Los Bafios, Luzon, Philip- 
pine Islands. 
Bezzi recorded the type as being in Profes- 
sor C. F. Baker’s collection (University of the 
Philippines). A specimen in the U. S. Na- 
tional Museum labeled "Mt. Makiling, Lu- 
zon, Baker, 1497” is evidently a cotype. 
The many thousands of specimens which 
have been studied from Malaya have shown 
considerable variation in the width of the 
costal band and in the intensity of the dis- 
coloration of the front and middle coxae and 
the anterior portion of the propleura. So much 
intergradation is evident that it does not ap- 
pear practical to attempt to separate occipitalis 
from the typical dorsalis in this region. The 
data recorded under dorsalis will also pertain 
to the variety occipitalis. In most of the series 
examined, specimens fitting both forms were 
present. 
Dacus {Strumeta) dorsaloides n. sp. 
Fig. 9 
This species is very close to D. dorsalis 
Hendel and to D. cognatus n. sp. and is best 
differentiated by the characteristics of the fe- 
male ovipositor. The bright-yellow postsu- 
tural vittae are similar to those of cognatus, 
they are narrowed behind and do not reach 
the inner supraalar bristles, but the area of the 
mesonotum outside the vittae is rufous or 
brownish yellow, not black as in the related 
species. The tibiae are also discolored with 
brown as in cognatus, but the humeri are bor- 
dered by rufous, not black. It differs from 
both dorsalis and cognatus by having the apical 
third of each femur discolored with brown. 
The females are readily distinguished from 
dorsalis by the very short, stubby ovipositor. 
They can be grouped with cognatus by the 
comparatively broad, gradually tapered apex 
(cf. Figs. Ga, 9a). It differs from both dorsalis 
and cognatus (as well as all other Dacinae which 
we have studied) by having the basal segment 
of the ovipositor longer than the inversion 
membrane or the piercer. It is further dis- 
tinguished by the details of the ovipositor as 
given below. 
FEMALE. Head: The front is approximately 
twice as long as wide; there are two pairs of 
inferior frontal orbital bristles and one pair of 
superior frontal orbitals. No brown spots are 
present at the bases of the frontal bristles. 
Fig. 9. Dacus {Strumeta) dorsaloides n. sp. a. Extended 
ovipositor; b, apex of piercer. 
