Sequoia gigantea •— St. John AND Krauss 
343 
a short petiole, in cross 
section with 1 resin duct 
below the fibrovascular 
bundle 
stomata on 1 or 4 sides, 
with 2 lateral resin ducts 
(or none) 
Taxodium 
Sequoia (in former broad 
sense, including S. sem- 
pervirens and S. gigantea) 
Staminate flowers in elon- 
gate spikes or panicles, 
scales with 5-9 sporangia 
Staminate cones ovoid, 
terminal or axillary, scales 
with 2-5 sporangia 
Pistillate cones subglo- 
bose or obovoid, scales 
2-seeded, thick, coriace- 
ous, peltate, the apex a 
4-sided, often mucronate 
disc 
Pistillate cones oblong- 
ovoid, scales 3-12-seeded, 
woody, wedge-shaped, 
often mucronate 
Seeds with 3 thick wings, 
cotyledons 4-9, embryos 
2 
Seeds with 2 wings, coty- 
ledons 2-5, embryos 2-5 
Buds scaly 
Buds scaly or naked 
Branchlets of 2 kinds, 
those near apex persist- 
ent, those lower and lat- 
eral deciduous 
Branchlets of 1 or 2 kinds, 
persistent 
Short shoots persistent 
Short shoots deciduous 
We should probably mention the recently 
described genus Metasequoia with one living 
species and many fossil ones, since there is 
such a mass of writing on it. Its generic name 
would suggest a close relationship to Sequoia, 
but it has many different characters, and, 
significantly, the cone scales are decussate 
instead of spiral. It is not a close relative o f 
Sequoia. The generic differences are summar- 
ized by Chaney (1951: 180). 
After this review of the characters that have 
proved significant and acceptable for the sep- 
aration of other pairs of closely related genera, 
we re-examine Sequoia. It is clear to the writers 
that Sequoia and Sequoiadendron are true gen- 
era, distinguished by many more contrasting, 
fundamental characters than those that form 
the basis for separation of numerous others 
among the widely accepted genera in the 
Coniferae. 
Accepting, now, as genera, the two units 
compared, we review the nomenclature of the 
living species. 
SYNONOMY OF SEQUOIA 
Sequoia sempervirens (D. Don in Lamb.) EndL, 
Syn. Conif. :'98, 1847. 
Taxodium sempervirens D. Don in Lamb., 
Gen. Pinus 2: 24, 1824; also ed. 2, 2: 
107, pL 48, 1828. 
Schuhertia? sempervirens (D. Don in Lamb.) 
Spach, Hist. Nat. Veg. 11: 353, 1842. 
{Schuhertia is a nomen genericum reji- 
ciendum.) 
Sequoia gigantea EndL, Syn. Conif. 198, 
1847, not of Dene. 1854 which is Se- 
quoiadendron giganteum. 
Sequoia religiosa Presl, Bdhmische Gesell. 
Wiss., Abhandl. V, 6: 597, 1851; and 
reprinted as Epimel Bot. 237, 1851. 
Condylocarpus Salisb., in Lamb., Gen. Pinus, 
ed. minor 2: 120, 1832, published by D. 
Don in synonomy of Taxodium semper- 
virens D. Don in Lamb. 
Gigantabies taxifoUa]. Nelson, under pseu- 
donym Senilis, Pinac. 78, 1866. 
Sequoia taxi folia Kirwan, Pinac. 246, 1868. 
Steinhauera sempervirens (D. Don in Lamb.) 
Voss, Deut. Dendrol. GeselL, Mitt. 16 
(1907): 90, 1908, the name Steinhauera 
now being a nomen genericum reji- 
ciendum. 
The customary name of this tree remains 
unchanged. 
SYNONOMY OF SEQUOIADENDRON 
Sequoiadendron giganteum (Lindl.) Buchholz, 
Amer. Jour. Bot. 26: 536-538, 1939- 
WelUngtonia gigantea Lindl., Gard. Chron., 
819-820, 823, 1853: and Hooker’s Jour. 
Bot. & Kew Misc. 7: 26, 1855; not 
WelUngtonia Meisn. (1840) of the Sa- 
biaceae. 
Americas gigantea (Lindl.) Anon., Descrip- 
tion of the Great Tree, recently felled 
upon the Sierra Nevada, California, now 
placed for public exhibition, in the spa- 
cious racket court of the Union Club, 
No. 596 Broadway, adjoining the Metro- 
politan Hotel, New York, p. 6-7, 1854. 
