18 
PACIFIC SCIENCE, VoL XII, January, 1958 
cited, as I plan to publish an annotated catalog 
of the flora of Micronesia, in which all speci- 
mens seen will be cited. Neither a description 
of the island nor of its vegetation are included 
as these will be given in a paper on the fossil 
plants (Fosberg and Corwin, 1958). After 
each species the workers who have recorded 
or collected it on Pagan are designated by the 
following symbols in bold-faced type: 
Marche 
M 
Fritz 
Fr 
Kanehira 
K 
Hospkawa 
H 
Anderson 
A 
Fosberg 
Fo 
Bonham 
B 
Corwin 
C 
The nomenclature is that presently accepted 
by me and not necessarily that under which 
the original reports were made. 
The indigenous flora is almost entirely an 
attenuation of that of the Marianas group to 
the south, some of the species extending 
northward to the Bonin Islands. Trema argen- 
tea and My op or um boninense may possibly be 
plants that have spread from the Bonin Is- 
lands southward in prehistoric time, though 
a spread in the opposite direction is entirely 
possible, as both species are known from 
Saipan. Fimbristylis urakasiana has previously 
been regarded as endemic on Uracas, the 
northernmost of the Marianas. It is very likely 
present at least on all of the recent volcanic 
islands of the group, but they have not been 
adequately enough studied and collected to 
detect all plants present, especially in such 
groups as Fimbristylis . Of the species and 
varieties, both recent and fossil, 20 are prob- 
ably endemic to the Marianas and 3 more to 
the Mariana, Volcano, and Bonin islands. 
The remainder are widespread species, mainly 
of lowlands and strands of the Indo-Pacific 
area, a few of them almost pantropic. These 
figures are probably valuable only as a general 
indication of the distributional relations on a 
rather young volcanic island. When more 
complete collections have been made, both 
on Pagan and on the other northern Marianas, 
more reliable and significant conclusions may 
be reached as to the origin of the flora of this 
area. 
SYSTEMATIC LIST OF SPECIES 
* Indicates recently introduced species; f in- 
dicates species believed to be of aboriginal in- 
troduction. 
Gleichenia linearis (Burm. f.) C.B.C1. Fo 
Cheilanthes tenuifolia (Burm.) Sw. H, Fo 
Acrostichum aureum L. H, Fo 
Asplenium laserpitiifolium Lam. FI 
Asplenium nidus L. A 
Asplenium unilaterale Lam. H 
Davallia solida (Forst. f.) Sw. H, A 
Dryopteris unita (L.) O. Ktze. Fo, B 
"Lastrea” torresiana (Gaud.) Moore A, Fo 
Nephrolepis hirsutula (Forst. f.) Presl H, 
A, Fo, B 
Polypodium scolopendria Burm. f. H, Fo, 
B 
Pteris quadriaurita Retz. H, Fo 
Sphenomeris chinensis (L.) Maxon Fo 
Psilotum nudum (L.) Griseb. H 
Freycinetia mariannensis Merr. H, A 
Pandanus dubius Spr. H 
Pandanus tectorius Park. K, H, A, B 
*Cenchrus brownii R. & S. A 
*Cenchrus echinatus L. A 
*Chloris inflata Link B 
Chrysopogon aciculatus (Retz.) Trin. A, 
Fo, B 
*Cynodon dactylon (L.) Pers. A, B 
*Dactyloctenium aegyptium (L.) Willd. H, 
A, Fo, B 
*Digitaria ciliaris (Retz.) Koel. Fo 
*Eleusine indica (L.) Gaertn. A, Fo 
*Eragrostis amabilis (L.) W. & A. H, A, Fo 
Heteropogon contortus (L.) Beauv. A, Fo 
Miscanthus floridulus (Labill.) Warb. H, K, 
A, B 
Panicum ambiguum Trin. A, Fo 
*Paspalum conjugatum Berg. Fo 
Paspalum orbiculare Forst. f. A, Fo 
*Pennisetum purpureum Schumach Fo 
Sporobolus virginicus (L.) Kunth A, B 
Thuarea involuta (Forst. f.) R. & S. H 
*Zea mays L. Fr 
