322 
PACIFIC SCIENCE, Vol. XII, October, 1958 
Fig. 4. Doliolina undulata Tokioka and Berner 1958, 
alimentary organs and gonads, from left side. 
very little undulation. In these cases the ven- 
tral part of the body, just posterior to muscle 
V, is distended by the testis. The appearance 
of the testis and its relative position against 
the intestinal loop are fundamentally the same 
as in specimens with the undulating testis. 
Subdivisions of the genus Doliolina 
With the descriptions of D. undulata Toki- 
oka and Berner (1958) and here of D. obscura 
and D. separata , we find two types of muscle 
pattern within the genus Doliolina . These pat- 
terns, one in which all of the muscles form 
complete loops and one in which the seventh 
muscle is interrupted ventrally, distinctly di- 
vide the genus into two groups. On this basis 
we have established the following subdivision 
of the genus: 
1) Doliolina perfecta: The muscle bands of 
this group all form complete bands. The fol- 
lowing species are included: indica (Neumann 
1906), intermedia (Neumann 1906), resistible 
(Neumann 1913), mulleri (Krohn 1852), 
krohni (Herdmann 1888), and sigmoides Gar- 
stang 1933. 
'2) Doliolina imperfecta: Within this group 
the seventh muscle band is always interrupted 
at the mid-ventral line. The species in this 
group are known only from the eastern Pa- 
cific Ocean, and include: undulata Tokioka 
and Berner 1958, separata n. sp., and obscura 
n. sp. 
The shape and position of the gonad and 
the arrangement of the gill septum vary con- 
siderably within the two groups, and in some 
cases convergence of these features may be 
seen. We thus hesitate to treat them as sub- 
genera of Doliolina but feel the problem may 
better be settled by more extensive collections 
and further critical examination of the re- 
spective species. 
The discovery of three species of doliolids, 
in addition to the one previously known 
{Dolioletta mirabilis) , in which a body muscle 
is interrupted gives support to the argument 
for the abandonment of the ordinal name 
Cyclomyaria for the doliolids, in favor of the 
name Doliolida. This change has been gen- 
erally accepted since Garstang’s review (1933). 
Doliopsoides horizoni Tokioka and Berner 
1958 
Figs. 5A, B 
Many additional specimens of D. horizoni 
have been found in the material from the 
Shellback Expedition. In some of these the 
alimentary organs are better preserved than 
in any of the previous specimens. Close ex- 
amination of these organs reveals that the 
stomach is provided with several fingerlike 
protuberances . In one especially well-preserved 
specimen, four pairs of these protuberances 
were found on the ventral, posterior portion 
of the cardiac stomach and two pairs on the 
ventral, posterior portion of the pyloric 
stomach. The number of protuberances is not 
constant among the specimens. 
