54 
PACIFIC SCIENCE, Vol XIX, January 1965 
Collections from wild: 
P. candelabrum Beauv. ...Akofokyir, 
Cape Coast, Ghana 
Freycinetia 
Collections from wild: 
F. arborea Gaud Poamoho Trail, 
Oahu, Hawaii 
F. mariannensis Merr.. .Stone 42 17, Guam 
Slide collection, Jodrell Laboratory, Royal 
Botanic Gardens, Kew: 
F. banksii Cunn. 
F. javanica Bl. 
F. lucens Ridl. 
Slide collection, Professor V. I. Cheadle, at 
University of California, Davis, California: 
F. excelsa F. Muell CA. 216 and 230 
Atherton and Cairns, Australia 
Sararanga 
Dried specimen from Herbarium, Royal 
Botanic Gardens, Kew: 
S. philippinensis Merr Ramos and 
Edano 48047, Philippines 
Collection from wild: 
S. sinuosa Hemsl G. F. C, Dennis 340, 
British Solomon Islands 
ACKNOWLEDGMENTS 
Technical assistants employed during this 
study were financed by a grant from the Ameri- 
can Philosophical Society. I wish to thank Mr. 
J. B. Hall for material of Pandanus candelabrum, 
Mr. G. C. Dennis for material of Sararanga sinu- 
osa? Mr. L. L. Forman for material of Sararanga 
philippinensis, Dr. C. R. Metcalfe and Dr. V. I. 
Cheadle for access to slide collections in their 
respective charge, Dr. Murray Gaskin for per- 
mitting unlimited access to plants cultivated at 
the U. S. Dept, of Agriculture Experiment Sta- 
tion, Old Cutler Road, Miami, and Professor 
T. M. Harris, F.R.S., for several helpful sugges- 
tions. I owe a particular debt to Dr. B. C. Stone, 
College of Agana, Guam, who originally made 
this study feasible and for his subsequent interest. 
REFERENCES 
Florin, R. 1931. Morphologic und Epidermis- 
struktur der Assimilationsorgane bei den rez- 
2 It is hoped to describe in detail the anatomy of 
Sararanga in a future account based on Mr. Dennis’s 
collections. 
enten Koniferen. K. Svensk. Vet. Handl. Ser. 
3, 10(1) :588. 
Kofler, Johanna. 1918. Der Dimorphismus 
der Spaltoffnungen bei Pandanus. Ost. Bot. 
Zeit. 67:186-196. 
Metcalfe, C. R. I960. Anatomy of the Mono- 
cotyledons, Vol. I. Gramineae. Oxford. 
— 1961. The anatomical approach to sys- 
tematics. Recent Advances in Botany. Toronto. 
Pp. 146-150. 
Pfitzer, E. 1870. Beitrage zur Kenntniss der 
Hautgewebe der Pflanzen. Jb. wiss. Bot. 7: 
532-587. 
St. John, H. I960. Revision of the genus Pan- 
danus Stickman. Part I. Key to the Sections. 
Pacif. Sci. 16:226. 
Schoute, J. C. 1903. Die Stammesbildung der 
Monokotylen. Flora, Jena 92:32-48. 
1906. Uber die Verastelung bei Mono- 
kotylen Baumen 1. Die Verastelung von Pan- 
danus. Ann. Jard. Bot. Buitenz. 20, ser. 2, 
5:53-87. 
— 1907. Uber die Verdickungsweise des 
Stammes von Pandanus. Ann. Jard. Bot. Bu- 
itenz. 21, ser. 2, 6: 115-137. 
Schumann, K. 1897. Die Verzweigung der 
Pandanaceen. Bot. Jahrber. 23:559-572. 
Solereder, H., and F. J. Meyer. 1933. Pandana- 
ceae. In: Systematische Anatomic der Mono- 
kotyledonen. Heft. 1:28-50. 
Solla, R. F. 1884. Contribuzione alio studio 
degli stomi delle Pandanee. Nuov. G. bot. ital. 
16 : 172 - 182 . 
Stebbins, G. L., and G. S. Khush. 1961. Varia- 
tion in the organization of the stomatal com- 
plex in the leaf epidermis of monocotyledons 
and its bearing on their phylogeny. Amer. J. 
Bot. 48:51-59. 
Tomlinson, P. B. 1961. Anatomy of the Mono- 
cotyledons, II. Palmae fed. by C. R. Metcalfe], 
Oxford. 
1964. Stem structure in arborescent 
monocotyledons. In: Formation of Wood in 
Forest Trees, pp. 65-86 [ed. by M. H. Zim- 
mermann]. New York. 
van Steenis, C. C. G. J. 1954. Pandanus in 
Malaysian vegetation types. Flora Malesiana 
I, 4( 1 ) : 3—12. 
Warburg, O. 1900. Pandanaceae in Das Pflan- 
zenreich. 4(a) : 97. 
