78 
PACIFIC SCIENCE, VoL XIX, January 1965 
fusiform, 0.2-0.26 mm long, 6 0-90/x wide; bi- 
polar filament 0.25-0.32 mm long. Vagina open- 
ing midventrally at a distance of 0.82-1.55 mm 
from head end, provided with a bulb of circular 
muscles near its comparatively wide, circular 
aperture; paired accessory vaginal duct aris- 
ing sideways from base of vagina, running 
transversely and then turning backward to join 
transverse commissure between anastomosing 
anteriormost vitelline ducts of two sides; median 
vaginal duct proper can be traced further back 
of this commissure, but its posterior termination 
is unknown. Vitellaria commencing in lateral 
fields at level of vaginal pore, confluent in post- 
ovarian region; vitelline reservoir Y-shaped with 
its stem between ovary and left intestinal limb. 
DISCUSSION : This genus differs from the most 
closely related Pseudopisthogyne in the body 
being distinctly constricted posterior to the 
widest midregion, in the ovary being widely sep- 
arated from the testes by this constricted area, 
and in the caudal appendage being reduced to a 
conical process bearing only one pair of hooks. 
That the lateral sclerites of the larger clamps do 
not show a sigmoid curve in contrast with those 
of Pseudopisthogyne or Pseudopisthogynopsis is 
also worth noting. 
Allopseudopisthogyne n. g. 
GENERIC DIAGNOSIS: Discocotylidae, Opis- 
thogyninae: Body elongate, asymmetrical, con- 
stricted a little posterior to widest midregion, 
without oblique body ridges posteriorly. Opis- 
thohaptor asymmetrical, with four large subses- 
sile clamps on side of genital pore and four 
smaller pedunculate clamps on other side; larger 
clamp with asymmetrical basal and lateral scle- 
rites, latter sclerites not showing sigmoid curve; 
smaller clamp with exactly symmetrical lateral 
sclerites. Caudal appendage reduced to conical 
process, with only one pair of hooks, attached to 
posterior end of body nearer to small clamp than 
to large clamp. Head blunt-pointed, with apical 
organs. Esophagus bifurcating at level of vaginal 
pore; intestinal limbs with side branches, not 
accompanied by vitellaria in constricted part of 
body, not confluent posteriorly. Testes very nu- 
merous, widely separated from ovary by body 
constriction. Pars prostatica, ejaculatory duct, 
cirrus pouch and genital atrium well differen- 
tiated just as in Pseudopisthogyne and Pseu- 
dopisthogynopsis. Cirrus unarmed. Common 
genital pore in esophageal zone, ventrosubmar- 
ginal. Ovary turned back on itself in posterior 
third of body, with both ends directed back- 
wards. Eggs fusiform, with bipolar filaments. 
Vagina provided with sphincter, opening mid- 
ventrally far posterior to common genital pore, 
at intestinal bifurcation; paired accessory vagi- 
nal ducts connected with transverse commissures 
between anteriormost anastomosing vitelline 
ducts of two sides; median vaginal duct proper 
rudimentary. Vitellaria co-extensive with intes- 
tine except at constricted part of body, confluent 
posterior to ovary; vitelline reservoir Y-shaped, 
sinistral to ovary; anterior vitelline ducts con- 
taining sperm. Gill parasties of marine teleosts. 
type SPECIES: A. constricta n. sp., on Lepi- 
docybium flavobrunnemn; Hawaii. 
12. Pseudopisthogynopsis lepidocybii n. g., n. 
sp. 
Fig. 12 
HABITAT: Gill of Lepidocybium flavobrun- 
neum; Hawaii. 
holotype: U. S. Nat. Mus. Helm. Coll., S.Y. 
No. 12. 
description (based on 50 whole mounts): 
Body elongate, 15-13 mm long, 0.55-1.4 mm 
wide just in front of opisthohaptor, tapering 
anteriorly, symmetrical except for posterior end, 
to which four larger subsessile clamps are at- 
tached on the side of the genital pore and four 
smaller pedunculate clamps on the other side, 
without hook-bearing appendage between two 
posteriormost clamps. Skeleton of small clamp 
100-170/x wide, consisting of two pairs of sym- 
metrical or slightly asymmetrical, unjointed, 
lateral sclerites, a median spring with bifid ends 
of which the shorter prong has a V-shaped apical 
piece, a pair of accessory sclerites and a pair of 
short basal sclerites; skeleton of larger clamp 
280-420 g wide at base, consisting of two pairs 
of distinctly asymmetrical, unjointed, lateral scle- 
rites, a median spring with bifid ends, a pair of 
accessory sclerites and a pair of basal sclerites. 
On the tip of the shorter, fenestrated prong of 
the median spring is set a V-shaped, more weakly 
sclerotized, apical piece. The capsule of the larger 
clamp is more muscular than that of the smaller 
